Re-description of the genus Heterobiantes and its type species H. geniculatus (Opiliones: Laniatores: Epedanidae) with additional genital and external morphology data Author Tan, Desmond Kai Teck 0009-0003-2388-7214 Science Unit, Lingnan University, Hong Kong, China & desmondtan @ ln. hk; https: // orcid. org / 0009 - 0003 - 2388 - 7214 desmondtan@ln.hk Author Fong, Jonathan J. 0000-0002-3648-6730 Science Unit, Lingnan University, Hong Kong, China & jonfong @ ln. edu. hk; https: // orcid. org / 0000 - 0002 - 3648 - 6730 jonfong@ln.edu.hk Author Zhang, Chao 0000-0003-1702-1206 Key Laboratory of Zoological Systematics and Application of Hebei Province, Institute of Life Sciences and Green Development, College of Life Sciences, Hebei University, Baoding, China & opiliones @ 163. com: https: // orcid. org / 0000 - 0003 - 1702 - 1206 text Zootaxa 2023 2023-10-13 5353 6 533 550 http://dx.doi.org/10.11646/zootaxa.5353.6.2 journal article 10.11646/zootaxa.5353.6.2 1175-5326 10012459 DD342940-639F-4054-B153-932C318DF73E Heterobiantes Roewer, 1912 Heterobiantes Roewer, 1912: 219 ; 1923: 191–192 , f. 217; Kury et al . 2020: 32–33. Epedanestus Roewer, 1938: 108 . Type species by monotypy: Epedanus geniculatus Pocock, 1903 . Synonymized by Kury et al . 2020: 32. Type Species: Epedanus geniculatus Pocock, 1903 , subsequent designation by Kury et al . 2020: 32. Diagnosis: Large-sized epedanines ( 5.94–6 mm ) with a long median spine on the ocularium. The body is trapezoidal in shape. Dorsal scutum distinctly segregated into four different areas (areas I–II fused). Two apparent yellow (sometimes pale green) spots are located on the posterior-lateral sides of prosoma. Areas I–II with a row of 8–10 granules, and often but not always with two that are positioned nearing the anterior margins of the area. Area V and all free tergites are lined with hair-tipped tubercles. Proximal segment of chelicerae strong and clavate, distally clustered with granules and hair-tipped tubercles. Pedipalps elongated, trochanter dorsally armed with one tubercle while femurs are slightly arcuate, armed with 3–5 small tubercles dorsally and 8–9 strong tubercles ventrally ( Figs 3 B-C). A disto-ental tubercle can also be found on the femur. Tibia and tarsus heavily armed with long spines on both ventral sides, which form a ‘basket claw’ ( Wolff et al . 2016 ) ( Figs 3A–C ). All distitarsi with three segments. Penis elongated and the shaft widens distally, armed with a lower transverse row of 7–10 strong, curved spines. The upper, subdistal section is protected by 10–15 similarly curved spines, along with three more pairs of ventral spines of glans. The positioning of the spines together resembles the shape of a crown, and is of resemblance to other Epedanidae members (e.g., K. insulanus ). Total length of the penis is 2.4–2.6 mm . Distribution: China ( Hong Kong ).