New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae) Author Bocek, Matej Author Adamkova, Kristyna text Zootaxa 2019 2019-06-25 4623 2 341 350 journal article 26407 10.11646/zootaxa.4623.2.8 db2b3bd4-2e27-4918-8fff-226b303c65b9 1175-5326 3255499 EEDC9982-ED9C-40BD-8584-CDADA572DDE Eniclases Waterhouse, 1879 Eniclases Waterhouse, 1879: 66 . Type species. Lycus luteolus Waterhouse, 1878 (by original designation). = Trichalolus Pic, 1923: 36 , hors texte; Bocak & Bocakova, 1991 . Type species. T. apertus Pic, 1923: 36 (by monotypy). Diagnosis. Eniclases shares shortened primary costa 1 with other trichaline genera and differs in unique V-shaped pronotal carinae ( Figs 8–9 ), secondary elytral costae are always present. Male genitalia have the characteristic cup-shaped phallic apex ( Figs 11–12 ) similar to those of Schizotrichalus but unlike the latter, the phallobase has an extensive membranous sac ( Figs 11–12 ). For detailed Eniclases redescription see Bocak & Bocakova (1991) . Distribution and diversity. The highest diversity of Eniclases is known from New Guinea . Altogether, 37 formally described species are known from the island and only Eniclases moluccanus has been reported from Halmahera ( Bocak & Bocakova 1991 , Bocak 2002 , Bocek & Bocak 2016 ). Remark. The molecular analysis supports morphology-based sister relationships of Schizotrichalus and Eniclases . Moreover, the deeply rooted Moluccan species indicate that the ancestral range of Eniclases is located in the combined area of New Guinea and the northern Moluccas .