Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia Author Jirapatrasilp, Parin https://orcid.org/0000-0002-5591-6724 Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Sutcharit, Chirasak Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand jirasak4@yahoo.com Author Panha, Somsak Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand somsak.pan@chula.ac.th text ZooKeys 2022 2022-08-25 1119 1 115 http://dx.doi.org/10.3897/zookeys.1119.85400 journal article http://dx.doi.org/10.3897/zookeys.1119.85400 1313-2970-1119-1 A3BE91C6B79344E1A886A803BF104D8B F158A7C5261D52B69288A62F3C777CAF Pupina godwinausteni Jirapatrasilp sp. nov. Figs 10C , 28Q, R , 32E, F , 33B, C Type material. Holotype CUMZ 12090/1 (Figs 10C , 28Q , 33B ), 5 June 2017, coll. C. Sutcharit, R. Srisonchai, A. Pholyotha. Measurement: shell height 8.8 mm, shell width 6.8 mm and 5 whorls. Paratypes CUMZ 12090/2-26 (24 shells and 1 specimen in ethanol; Figs 28R , 32E , 33C ) and NHMUK 20210335 (3 shells), same data as holotype; CUMZ 12091 (20 shells and 24 specimens in ethanol; Fig. 32F ) from the type locality, 5 Dec. 2020, coll. P. Jirapatrasilp, C. Sutcharit, A. Pholyotha. Type locality. Khao Wong Cave, Ban Rai District, Uthai Thani Province, Thailand, 15°01'52.6"N , 99°27'23.3"E , 246 m amsl. Other material examined. CUMZ 12092 (2 shells) from Tham Namthip Bureau of Monks , Lan Sak District , Uthai Thani Province , 28 July 2016 . CUMZ 12093 ( 1 specimen in ethanol) from Hup Pa Tat , Lan Sak District , Uthai Thani Province , 1 Oct. 2018 . Diagnosis. Shell globose; last whorl ca. 80% of shell height. Apertural lip very thickened and slightly expanded; with a furrow between inner and outer peristomes; inner peristome thickened, cord-like; apertural lip angled when observed from lateral view. Columellar tooth curved downward like an earlobe. Differential diagnosis. The globose shell shape of P. godwinausteni sp. nov. is most similar to P. siamensis , but P. godwinausteni sp. nov. differs from P. siamensis in having a larger shell, a more prominent parietal callus, a thicker apertural lip with a furrow between inner and outer peristomes, with an inner peristome thickened and cord-like, a longer posterior canal, a wider and more curved columellar tooth, and a more angled apertural lip when observed from lateral view. Description. Shell height 7.7-9.5 mm; shell width 5.5-7.0 mm. Shell globose, solid, semi-transparent, brown, devoid of prominent sculpture on glazed smooth surface. Apex obtuse. Growth lines on shell surface inconspicuous. Whorls 5, last whorl large (ca. 80% of shell height) and bulging. Spire angle ca. 90°, somewhat extended. Sutures slightly impressed, but shallow. Aperture circular; lip highly thickened (ca. 0.4-0.5 mm wide and 0.5-0.6 mm thick) with darker colour, slightly expanded; apertural lip when observed from lateral view angled. Apertural lip with a furrow between inner and outer peristomes, with inner peristome thickened and cord-like. Parietal callus thickened with darker colour. Peristome interrupted by two canals; posterior canal ca. 1.0-1.2 mm long, 0.6 mm at its widest, curved outward and bulging at the outer margin; anterior canal slit-like, as long as apertural lip width. Parietal tooth thick; columellar tooth curved downward like an earlobe (ca. 2.2 mm long, 1.2 mm wide and 0.5 mm thick), covering anterior canal. Umbilicus closed. Operculum round, yellowish, and transparent corneous with uneven edge. Etymology. The specific epithet is dedicated to H.H. Godwin-Austen, a British malacologist, who prominently contributed to malacological studies in South and Southeast Asia. Distribution. This new species is found in Uthai Thani Province, Thailand.