Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae) Author Machado, Renato Jose Pires Author Oswald, John David text Zootaxa 2020 2020-06-16 4796 1 1 322 journal article 10.11646/zootaxa.4796.1.1 1175-5326 3896810 66DD1FEB-6BDE-4AEB-8A7B-96594371E9C5 Riekoleon convergens New, 1985 ( Figs. 151–153 ) Riekoleon convergens New, 1985:75 (OD); New 1996:93 (cat); Stange 2004:108 (cat); Oswald 2018 (cat). Diagnosis. Legs elongate; wings narrow and with numerous brown marks; tibial spurs reaching apex of T2; body mostly pale; pronotum with some long white setae; male ectoproct with a tubular posteroventral lobe. Description. Lengths: forewing: 19–21 mm ; hind wing: 18–19 mm . Head ( Figs. 151 a–b): Labrum pale, set with a line of elongate setae. Clypeus pale with a thin central longitudinal dark brown line, margin between clypeus and frons dark brown; set with some elongate white setae. Frons ventral margin pale with a thin central longitudinal dark brown line, but remaining areas dark brown; covered with short white setae. Gena pale. Vertex entirely pale with some small irregular brown marks posteriorly in some specimens; covered with short black setae. Ocular setae absent. Antennae clubbed; elongate, about twice longer than pronotum; distance between antennae wider than scape width; scape and pedicel mostly dark brown, most flagellomeres brown basally and pale distally, apex entirely dark brown; torular membrane pale; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae, except for scape and pedicel with short white setae. Mandible pale, with tip darker. Palpi , maxillary and labial pale with distal segments brown, and some irregular brown marks in some specimens; apical labial palpomere fusiform, palpimacula opening ovalshaped, located medially. Thorax ( Fig. 151b ): Pronotum longer than wide; posterior margin wider than anterior; subapical furrow present; mostly pale except for lateral margins up to furrow dark brown, and a thin longitudinal dark brown line medially in some specimens; set with few short black setae and some long white setae. Mesonotum colouration mostly pale with brown areas as follows: two rounded marks on posterior margin of prescutum, and two large rounded lateral marks at scutum; covered with short black setae, and few long white setae. Metanotum dark brown laterally with a broad longitudinal pale line medially, covered with few short black setae. Pterothoracic pleura mostly dark brown with irregular pale marks ventrally in some specimens; covered with long white setae; Miller’s organ present. Wings ( Fig. 151c ): Rather narrow with tip acute; anterior Banksian line present in both wings, posterior absent; veins mostly dark brown, but some veins intercalated with white spots, beset with short black setae. Male pilula axillaris present. Forewing membrane mostly hyaline with irregular brown marks, mainly near pterostigma (apex white), posterior margin, and around basal crossveins; CuA fork located near RP origin; three presectoral crossveins; subcostal veinlets simple; posterior area narrower than prefork area. Hind wing membrane mostly hyaline except for a small brown mark on rhegmal area and other two brown marks around pterostigma (white); MP fork located between RP origin and first fork, subcostal veinlets simple; one presectoral crossvein. Legs ( Figs. 151 a–b): All pairs of legs , femur elongate (> 3.5x length of coxa); tibia slightly longer than femur and more than twice longer than tarsi; tibial spurs surpassing T2 apex; T2, T3 and T4 about same size, T1 longer than T2, and T5 twice longer than T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with black setae, and scattered long ones. Proleg sense hair absent; tibia with antennal cleaning setae ventroapically; coxa and trochanter pale, femur mostly pale but apex and anterior surface dark brown, tibia pale with scattered dark brown areas; tarsi dark brown. Mesoleg with colour pattern similar to proleg, except for femur pale with apex and two thin lines on dorsal surface dark brown. Metaleg with femur and tibia slightly longer than anterior legs; colour pattern similar to mesoleg, except for tibia paler. Abdomen: Mostly pale with scattered dark brown marks, mainly on distal margin, but mostly dark brown in some specimens. Beset with short black setae. Male Terminalia ( Figs. 152 a–c): Ectoproct with posteroventral margin extended in lateral view; set with elongate black setae. 9 th sternite short, with posterior margin elongate medially in ventral view; covered with elongate black setae. Gonarcus large, thin and “C” shaped in lateral view; in posterior view with medial region broader. Mediuncus membranous. Paramere elongate longitudinally, with tip acute, base divided into two lobes and curving upwards in lateral and posterior view. Dorsal region of paramere and gonarcus connected by a membrane. Female Terminalia ( Figs. 152 d–e): Ectoproct posterior margin rounded covered with thin elongate setae. Lateral gonapophyses rounded, much smaller than ectoproct, set with some elongate setae, and few short and thickened setae ventrally. 7 th sternite small, with distal margin rounded in ventral view; covered with long black setae. Pregenital plate weakly sclerotized, small, a transverse bar with anterior margin straight, posterior margin with a medial rounded expansion in ventral view. Posterior gonapophyses broad, long, covered with many long black setae and many cavisetae dorsoapically. 9 th tergite with a broad membranous digitiform process. Anterior gonapophyses as a large plate covered with long setae. Ventral membrane gonapophyseal plates absent. FIGURE 151. Riekoleon convergens : a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal (ends of legs showing through wings at base). FIGURE 152. Riekoleon convergens : male : a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female : d) terminalia, lateral; e) terminalia, posteroventral (note pregenital plate hidden beneath sternite). Scale bars = 0.5 mm. Distribution ( Fig. 153 ). Australia : QLD. Known only from northeastern Queensland . Adult activity period. Records for November to February. Biology. Unknown, larva unknown. Name-bearing type . Riekoleon convergens : Holotype (by original designation), female, ANIC, examined. Type locality: Australia , Queensland , near Tully (town), Mission Beach. From original description: “ Holotype , ♀ , Queensland , Mission Beach, near Tully, 5–23.xi.1967 , R. Dobson ( ANIC )”. Condition: good; terminalia dissected. Additional material examined ( 3♂ , 8♀ ). AUSTRALIA : Queensland : Bald Hills Station : 30 km N of Cooktown , 11.ii.1982 , M.S. & B.J. Moulds ( 1♀ , AMSA ) ; Cape York : Silver Plains Homestead, 28.i.1962 , J.L. Wassell ( 1♀ PT , ANIC ) ; Crystal Creek : 22 miles SSE of Ingham , 18 o 58’S– 146 o 16’E , 9.xii.1968 , Britton & Misko ( 1♀ PT , ANIC ) ; Etty Bay : near Innisfail , 23.ii.1982 , M.S. & B.J. Moulds ( 1♀ , AMSA ) ; Half Tide : near McKay , xi.1965 ( 1♀ , QMBA ) ; Isabela Falls : NW Cooktown , 15 o 18’02’’S– 145 o 00’13’’E , 11.i.2004 , Moulds & Cowan ( 1♀ , AMSA ) ; 5 miles S of Kuranda , Speewah Road , 11.i.1967 , D. K. McAlpine and G. Holloway ( 1♂ , PT , AMSA ) ; McIlwraith Range : NE of Coen , 13 o 43.247’S– 143 o 19.458’E , 508m , 7.i.2007 , K. Hill , D. Marshall & M. Moulds ( 1♂ , AMSA ) ; Meringa : 27.ii.1926 ( 1♀ , QMBA ) ; Station Creek : 15 km NW of Mount Molloy , 26.xii.1987 , M.S. & B.J. Moulds ( 1♂ , FSCA , 1♀ , AMSA ) . Comments. See Comments for R. furcatus .