The wasp genus Sphex in Sub-Saharan Africa (Hymenoptera: Sphecidae) Author Dörfel, Thorleif H. 11B5C093-23D5-417C-AB64-65764FC2AF05 Museum für Naturkunde, Invalidenstrasse 43, 10115 Berlin, Germany. thorleif.doerfel@mfn.berlin Author Ohl, Michael 878259F2-C3C6-4264-B04A-C397E01E5C8E Museum für Naturkunde, Invalidenstrasse 43, 10115 Berlin, Germany. michael.ohl@mfn.berlin text European Journal of Taxonomy 2022 2022-02-23 796 1 1 170 http://dx.doi.org/10.5852/ejt.2022.796.1665 journal article 55605 10.5852/ejt.2022.796.1665 9c3fa32d-4320-4170-83e3-a5c045a4ba68 2118-9773 6299440 76C5C9C4-C6C1-4EDC-8FF8-9828A6EF2040 Sphex stadelmanni rufus subsp. nov. urn:lsid:zoobank.org:act: 4873E4CA-248F-4293-852F-0144E8E2E1D9 Figs 96–97, 99 (purple) Differential diagnosis Females of S. stadelmanni rufus subsp. nov. ( Fig. 96 ) are easy to recognize within the bohemanni group because of their predominantly ferruginous clypeus, femora and tibiae. These areas are largely black in other species, with at most the free clypeal margin being ferruginous. In males ( Fig. 97 ), the ferruginous color on the clypeus is less extensive and sometimes barely visible, but the leg coloration is concurrent with that of the female and often sufficient for identification as well. Another character that differs from S. stadelmanni stadelmanni is the color of the appressed facial setae, which is silvery-white in the nominate species, but brassy in this subspecies. Both are identical in genital structure, and the shape of their penis valvae ( Figs 21–22 ) reliably separates them from other species in the group. Etymology ‘ Rufus ’ is a Latin adjective meaning ‘red’, which refers to the conspicuously-colored legs and clypeus of this taxon. Material examined Holotype KENYA – Taita-Taveta County • 1 ♂ ; Wundanyi ; [ 3°23ʹ54ʺ S , 38°21ʹ37ʺ E ]; 5–10 Apr. 1997 ; Ma. Halada leg.; THD-014-OOLM ; GenBank CO1 gene: MW538562 ; OÖLM . Paratypes KENYA – Taita-Taveta County • 5 ♂♂ , 1 ♀ ; same collection data as for holotype; OÖLM • 3 ♂♂ , 2 ♀♀ ; same collection data as for holotype but 18–22 Mar. 1997 ; OÖLM . TANZANIA • 1 ♀ ; “German East Africa”; F. Fischer leg.; ZMB . – Kilimanjaro Region • 1 ♂ , 2 ♀♀ ; “ Kilimanjaro ”; [ 3°04ʹ33ʺ S , 37°21ʹ12ʺ E ]; Jan. 1906 ; Schröder leg.; ZMB • 1 ♀ ; Gonja ; [ 4°16ʹ09.3ʺ S , 38°02ʹ54.9ʺ E ]; 10 Jan. 1906 ; Schröder leg.; ZMB . – Morogoro Region • 1 ♀ ; “ Usaramo , Rufidji , Khutusteppe ”; [ 7°47ʹ59.4ʺ S , 37°29ʹ13.1ʺ E ]; Oct.–Nov. 1896 ; Götze leg.; ZMB . – Tanga Region • 1 ♀ ; early 1904; A. Karasek leg.; ZMB • 1 ♂ , 1 ♀ ; Dec. 1905 ; Schröder leg.; ZMB • 1 ♀ ; “ East Usambara ”; [ 4°59ʹ54.1ʺ S , 38°41ʹ37.7ʺ E ]; F. Fischer leg.; ZMB • 1 ♀ ; Amani Research Institute ; [ 5°05ʹ S , 38°40ʹ E ]; 2 Mar. 1908 ; J. Vosseler leg.; ZMB • 1 ♀ ; Bumbuli ; [ 4°52ʹ S , 38°28ʹ E ]; 15 Dec. 1902 ; Meinhof leg.; ZMB . Description Female SIZE. 25.0– 31.5 mm . COLOR. Black except for the following, which are ferruginous: basal half of mandible, clypeus excepting dorsal and lateral margin, basal half of scape, trochanters, femora and tibiae. Wings uniformly fuscous, with purple-bluish iridescence. VESTITURE. Appressed setae on clypeus and paraocular area black with ferruginous-golden luster, on collar, scutum and propodeal enclosure black. Erect setae on clypeus, paraocular area, collar, scutum and propodeal enclosure black. Erect propodeal setae oriented posteriorly. Lower center of clypeus glabrous. Scutellum densely and coarsely pubescent. STRUCTURE. Free clypeal margin medially with two broad, indistinct processes, stepped above. Clypeus with inconspicuous indentation in lower center, longitudinal carina in upper center barely noticeable. Scutellum convex. Metanotum slightly raised, not notably bituberculate. 2 nd recurrent vein joins markedly proximal from interstitium between submarginal cells II and III. Propodeal enclosure without any notable ridges. Foretarsomere I 2.5–2.8 × length of antepenultimate spine. Petiole length 1.3–1.9 × its medial width. Male SIZE. 24.5–28.2 mm . COLOR. Black except for the following, which are ferruginous: basal half of mandible, longitudinal stripe medially on clypeus, basal half of scape, trochanters, femora and tibiae. Wings slightly fuscous, with violet iridescence. VESTITURE. Appressed setae on clypeus and paraocular area brassy, on collar, scutum and propodeal enclosure black, on posterior margin of propodeum silvery-golden. Erect setae on clypeus black, on paraocular area black intermixed with silvery ones, on collar, scutum and propodeal enclosure black, on posterior margin of propodeum silvery-golden. Erect propodeal setae oriented posteriorly. Free clypeal margin medially glabrous. Scutellum densely and coarsely pubescent. Sterna II–VII laterally with increasingly dense fringes of black setae, sternum VIII uniformly with shorter, erect, ferruginous-brown setae which are longer laterally and posteriorly. STRUCTURE. Free clypeal margin simple. Scutellum convex. Metanotum slightly raised, not bituberculate. 2 nd recurrent vein joins markedly proximal from interstitium between submarginal cells II and III. Posterior margin of metasomal tergum VII convex. Propodeal enclosure without any notable ridges. Posterior margin of metasomal sternum VII simple, of metasomal sternum VIII semicircular. Petiole length 1.9–2.1× its medial width. Flagellomeres IV–VI with broad placoids covering their entire length, flagellomere VII sometimes with moderately broad placoid covering approximately its basal half. Variation Unknown. Distribution Eastern Africa.