<document id="3B9E68613A6F8928CCF53BEFF95CCAAA" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69DA092E7FE46FA7FCBAA2F8B" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius heterosquamosus Heller 1910" docType="treatment" docVersion="10" lastPageNumber="189" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="189" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1D852C683628035C69BAE86C4D955CE5">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="A20804A6ABBB62082625F83388E69907">Telnov, Dmitry</mods:namePart>
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<mods:title id="D7D82CD7ABFEA553F2A368DB0644F331">European Journal of Taxonomy</mods:title>
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<mods:date id="CB304BED538BEA0389FC3BA84DC5C8E7">2020</mods:date>
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<mods:identifier id="5ACFD89D6D6DEDE3931DEB7882DBB1B0" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69DA092E7FE46FA7FCBAA2F8B" ID-DOI="http://doi.org/10.5281/zenodo.4344491" ID-GBIF-Taxon="168648106" ID-Zenodo-Dep="4344491" LSID="urn:lsid:plazi:treatment:1A3787F69DA092E7FE46FA7FCBAA2F8B" httpUri="http://treatment.plazi.org/id/1A3787F69DA092E7FE46FA7FCBAA2F8B" lastPageNumber="189" pageId="188" pageNumber="189">
<subSubSection id="DA84656B9DA092E7FE46FA7FCC092DBE" box="[469,1119,1473,1500]" pageId="188" pageNumber="189" type="nomenclature">
<paragraph id="922136E09DA092E7FE46FA7FCC092DBE" blockId="188.[469,1119,1473,1500]" box="[469,1119,1473,1500]" pageId="188" pageNumber="189">
<heading id="C969818C9DA092E7FE46FA7FCC092DBE" box="[469,1119,1473,1500]" centered="true" fontSize="11" level="2" pageId="188" pageNumber="189" reason="2">
<taxonomicName id="559E4D639DA092E7FE46FA7FCB802DB9" ID-CoL="7X3WT" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[469,982,1473,1500]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29DA092E7FE46FA7FCB6A2DBE" bold="true" box="[469,828,1473,1500]" italics="true" pageId="188" pageNumber="189">Maechidius heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119DA092E7FCD7FA7FCB802DB9" author="Heller K. M." box="[836,982,1473,1499]" pageId="188" pageNumber="189" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
comb. rest.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9DA092E7FF2EF9BFCB1F2E1C" pageId="188" pageNumber="189" type="reference_group">
<paragraph id="922136E09DA092E7FF2EF9BFCBBA2E7E" blockId="188.[189,1028,1537,1598]" box="[189,1004,1537,1564]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="B28E11CE9DA092E7FF2EF9BFCBBA2E7E" box="[189,1004,1537,1564]" pageId="188" pageNumber="189">
<treatmentCitation id="133F10F19DA092E7FF2EF9BFCAB62E79" author="Heller K. M." box="[189,736,1537,1563]" page="23" pageId="188" pageNumber="189" year="1910">
<taxonomicName id="559E4D639DA092E7FF2EF9BFCAB62E79" ID-CoL="7X3WT" authority="Heller, 1910: 23" authorityName="Heller" authorityPageNumber="23" authorityYear="1910" box="[189,736,1537,1563]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29DA092E7FF2EF9BFCA4E2E79" box="[189,536,1537,1563]" italics="true" pageId="188" pageNumber="189">Maechidius heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119DA092E7FD8CF9BFCAB62E79" author="Heller K. M." box="[543,736,1537,1563]" pageId="188" pageNumber="189" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910: 23</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, (original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="922136E09DA092E7FF2EF99DCC522E5C" blockId="188.[189,1028,1537,1598]" box="[189,1028,1571,1598]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="B28E11CE9DA092E7FF2EF99DCC522E5C" box="[189,1028,1571,1598]" pageId="188" pageNumber="189">
<treatmentCitation id="133F10F19DA092E7FF2EF99DCAE52E5C" author="Frey G." box="[189,691,1571,1598]" page="505" pageId="188" pageNumber="189" year="1969">
<taxonomicName id="559E4D639DA092E7FF2EF99DCAE52E5C" ID-CoL="7XWML" authority="Frey, 1969: 505" authorityName="Frey" authorityPageNumber="505" authorityYear="1969" box="[189,691,1571,1598]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29DA092E7FF2EF99DC9A22E5F" box="[189,500,1571,1597]" italics="true" pageId="188" pageNumber="189">Paramaechidius clypeatus</emphasis>
<bibRefCitation id="F60F4B119DA092E7FE68F99DCAE52E5C" author="Frey G." box="[507,691,1571,1598]" pageId="188" pageNumber="189" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969: 505</bibRefCitation>
</taxonomicName>
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(key and new combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="922136E09DA092E7FF2EF9DDCB1F2E1C" blockId="188.[189,841,1634,1662]" box="[189,841,1634,1662]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="B28E11CE9DA092E7FF2EF9DDCB1F2E1C" box="[189,841,1634,1662]" pageId="188" pageNumber="189">
<taxonomicName id="559E4D639DA092E7FF2EF9DDCA4E2E1F" ID-CoL="7X3WT" authorityName="Heller" authorityYear="1910" box="[189,536,1635,1661]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29DA092E7FF2EF9DDCA4E2E1F" box="[189,536,1635,1661]" italics="true" pageId="188" pageNumber="189">Maechidius heterosquamosus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="133F10F19DA092E7FDA6F9DDCAAA2E1F" author="Heller K. M." box="[565,764,1634,1661]" page="625" pageId="188" pageNumber="189" year="1914">
<bibRefCitation id="F60F4B119DA092E7FDA6F9DDCAAA2E1F" author="Heller K. M." box="[565,764,1634,1661]" pageId="188" pageNumber="189" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller 1914: 625</bibRefCitation>
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9DA092E7FF2EF91DCAAD2F6E" pageId="188" pageNumber="189" type="materials_examined">
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<heading id="C969818C9DA092E7FF2EF91DC93A2EDE" bold="true" box="[189,364,1698,1725]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<materialsCitation id="22F63CBD9DA092E7FF2EF91DC93A2EDE" ID-GBIF-Occurrence="2898878396" box="[189,364,1698,1725]" pageId="188" pageNumber="189" typeStatus="lectotype">
<emphasis id="A0EAEAF29DA092E7FF2EF91DC93A2EDE" bold="true" box="[189,364,1698,1725]" pageId="188" pageNumber="189">
<typeStatus id="4D2588429DA092E7FF2EF91DC8AC2EDF" box="[189,250,1699,1725]" pageId="188" pageNumber="189">Type</typeStatus>
material
</emphasis>
</materialsCitation>
</heading>
</paragraph>
<paragraph id="922136E09DA092E7FF2EF971CAAD2F6E" blockId="188.[189,798,1698,1804]" pageId="188" pageNumber="189">
<taxonomicName id="559E4D639DA092E7FF2EF971CA242E88" authority=": SNSD" authorityName="SNSD" box="[189,626,1743,1770]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29DA092E7FF2EF971CA4E2E8B" box="[189,536,1743,1769]" italics="true" pageId="188" pageNumber="189">Maechidius heterosquamosus</emphasis>
: 
<collectionCode id="F48FAE259DA092E7FDB4F96ECA242E88" box="[551,626,1744,1770]" collectionName="SNSD" pageId="188" pageNumber="189">SNSD</collectionCode>
</taxonomicName>
(
<specimenCount id="8498FD699DA092E7FD16F96ECACF2E88" box="[645,665,1744,1770]" pageId="188" pageNumber="189" type="female">♀</specimenCount>
<typeStatus id="4D2588429DA092E7FD32F971CB442E88" box="[673,786,1743,1770]" pageId="188" pageNumber="189" type="lectotype">lectotype</typeStatus>
). 
<taxonomicName id="559E4D639DA092E7FF2EF94FCA032F6E" authority=": BPBM" authorityName="BPBM" box="[189,597,1777,1804]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29DA092E7FF2EF94FC9A22F69" box="[189,500,1777,1803]" italics="true" pageId="188" pageNumber="189">Paramaechidius clypeatus</emphasis>
: 
<collectionCode id="F48FAE259DA092E7FD90F94CCA032F6E" box="[515,597,1778,1804]" collectionName="USA, Hawaii, Honolulu, Bernice P. Bishop Museum" pageId="188" pageNumber="189">BPBM</collectionCode>
</taxonomicName>
(
<specimenCount id="8498FD699DA092E7FDF5F94FCA2D2F69" box="[614,635,1777,1803]" pageId="188" pageNumber="189">♂</specimenCount>
<typeStatus id="4D2588429DA092E7FD10F94FCAB92F6E" box="[643,751,1777,1804]" pageId="188" pageNumber="189" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9DA092E7FF2EF88FCA312F1B" box="[189,615,1841,1913]" pageId="188" pageNumber="189" type="distribution">
<paragraph id="922136E09DA092E7FF2EF88FC9002F29" blockId="188.[189,342,1841,1867]" box="[189,342,1841,1867]" pageId="188" pageNumber="189">
<heading id="C969818C9DA092E7FF2EF88FC9002F29" bold="true" box="[189,342,1841,1867]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<emphasis id="A0EAEAF29DA092E7FF2EF88FC9002F29" bold="true" box="[189,342,1841,1867]" pageId="188" pageNumber="189">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09DA092E7FF2EF8E0CA312F1B" blockId="188.[189,615,1886,1913]" box="[189,615,1886,1913]" pageId="188" pageNumber="189">
North New 
<collectingCountry id="EA8976709DA092E7FED4F8E0C9CA2F1A" box="[327,412,1886,1912]" name="Guinea-Bissau" pageId="188" pageNumber="189">Guinea</collectingCountry>
(Torricelli Mts.).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9DA092E7FF2EF820CBAA2F8B" box="[189,1020,1950,2025]" pageId="188" pageNumber="189" type="discussion">
<paragraph id="922136E09DA092E7FF2EF820C9792FDA" blockId="188.[189,303,1950,1976]" box="[189,303,1950,1976]" pageId="188" pageNumber="189">
<heading id="C969818C9DA092E7FF2EF820C9792FDA" bold="true" box="[189,303,1950,1976]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<emphasis id="A0EAEAF29DA092E7FF2EF820C9792FDA" bold="true" box="[189,303,1950,1976]" pageId="188" pageNumber="189">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09DA092E7FF2EF870CBAA2F8B" blockId="188.[189,1020,1998,2025]" box="[189,1020,1998,2025]" pageId="188" pageNumber="189">
The male 
<typeStatus id="4D2588429DA092E7FEA1F871C9C12F8B" box="[306,407,1999,2025]" pageId="188" pageNumber="189" type="paratype">paratype</typeStatus>
from NHMB belongs to 
<taxonomicName id="559E4D639DA092E7FD2CF870CBCC2F8A" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[703,922,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="lapsus" status="sp. nov.">
<emphasis id="A0EAEAF29DA092E7FD2CF870CBCC2F8A" box="[703,922,1998,2024]" italics="true" pageId="188" pageNumber="189">Maechidius lapsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899DA092E7FC32F871CBAA2F8B" box="[929,1020,1999,2025]" pageId="188" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>