Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar
Author
Ünal, Mustafa
text
Zootaxa
2016
4206
1
1
223
journal article
37403
10.11646/zootaxa.4206.1.1
5c5bff7d-ea8c-4893-b03e-a8b5c5904514
1175-5326
208260
E1566C02-9987-4116-83AA-91D3D1DCF2FF
Nocaracris rubripes
(Motschulsky, 1846)
comb. rev.
(
Figs. 357
,
390
,
663–664
,
844
)
Eremoxenus rubripes
, de Motschulsky
in lit.: sensu Fischer von Waldheim 1846: 270.
Nocarodes rubripes
Motschulsky
: in Fischer von Waldheim 1846: 270.
Nocarodes rufipes
: Fischer von Waldheim 1846: 406
, pl. 31,
Fig. 3
, misspelling of
rubripes
.
Nocarodes rubripes
Motsch.
:
Fieber 1853
: 127
.
Nocarodes rufipes
:
Fieber 1853
: 127
, as syn. of
rubripes
.
Nocarodes cyanipes
Fischer de Waldheim, 1846
:
Kirby 1910
: 355
, partim;
I. Bolívar 1916
: 25
, partim.
Pamphagus (Nocarodes)cyanipes
Fisch. W.
:
Stål 1876
: 28
, partim.
Pamphagus (Nocarodes) rubripes
Fisch. W.
:
Stål 1876
: 28
, as syn. of
cyanipes
.
Nocarodes rubripes
Fischer de Waldheim, 1846
: Jacobson 1905: 200, 298, sensu
Bey-Bienko & Mistshenko 1951
;
I. Bolívar 1916
: 25
, as syn. of
cyanipes
.
Nocarodes rufipes
Fischer de Waldheim, 1846
:
I. Bolívar 1916
: 25
, as syn. of
cyanipes
.
Nocaracris rubripes
(F.-W.):
Uvarov 1928
: 150
;
Miram 1938
: 52
;
Tarbinsky 1940
: 34
, 214, 215;
Ramme 1951
: 411
, 427.
Nocaracris (Paranocaracris) rubripes
F.-W.:
Bey-Bienko 1964
: 251
.
Nocaracris tibialis
Ramme 1951
: 279
, 298, 427;
Avakyan 1968
: 100
, as syn. of
P. rubripes
;
Presa & Garcia 1983
: 17
;
Otte 1994
: 173
.
confirmed syn.
Paranocaracris rubripes
(Fischer de Waldhein, 1846)
:
Bey-Bienko & Mistshenko 1951
: 351
;
Karabağ 1958
: 122
;
Mirzayans 1959
: 16
;
Shumakov 1963
: 57
;
Avakyan 1968
: 100
;
Weidner 1969
: 160
;
Mistshenko 1972
: 52
; Stolyarov 1974: 481;
Presa & Garcia 1983
: 23
;
Heller
et al.
1998
: 61
.
Paranocaracris rubipes
(Fischer von Waldheim, 1846)
:
Hodjat 2012
: 267
;
Sevgili
et al.
2012
: 325
.
Paranocaracris rubripes rubripes
(Fischer de Waldheim, 1846)
:
Demirsoy 1973
: 424
, 425, partim;
Demirsoy 1977
: 81
, 82, partim;
Salman 1978
: 70
;
Erman & Salman 1990
: 112
;
Otte 1994
: 183
;
Stolyarov 1997
: 69
;
Mol
et al.
2014
: 137
.
Type
locality.
Armenia
(
Fischer von Waldheim
1846: 270).
Holotype
:
male (
NMW
).
Material
examined.
ARMENIA
:
Caucasus
,
1♂
(
Holotype
) (det.
Fieber
as
Nocarodes rubripes
) (det.
Br.
v.W. as
Nocarodes cyanipes
) (det.
Ramme
and
Cejchan
as
Nocaracris cyanipes
) (
NMW
)
;
Eriwan
,
Krasikoparan
,
1♀
(leg.
Christoph
) (det.
Br.
v.W. as
Nocarodes
spec.) (
NMW
);
Diližan
,
4.7.1926
,
1♀
(
Holotype
of
Nocaracris tibialis
) (leg.
Ižmailov
) (
MfN
); circ. lac. Gokča, prope Čubuchlu,
12.7.1927
,
2♂
,
4♀
;
Gokča
, pr.
Šordja
,
20.7.1927
,
1♂
;
Šordja
,
20.6.1928
,
1♂
,
4♀
;
Šordja
,
26.6.1928
,
3♂
;
Šordja
,
13.5.1930
,
6♀
;
Šordja
,
14.5.1930
,
3♂
,
5♀
;
Šordja
,
20.5.1930
,
1♂
,
2♀
;
Gokča
, pr.
Elenovka
,
21.8.1926
,
1♂
;
Elenovka
,
5.6.1927
,
3♀
;
Elenovka
,
14.6.1927
,
6♂
,
12♀
,
Elenovka
, 9
–
10.5.1930
, 8♂, 7♀;
Elenovka
,
3.7.1930
,
12♂
,
14♀
;
Gokča
, pr.
Täk-Agač
,
16.7.1927
,
13♂
,
12♀
,
1♀
nymph
;
Alagöz
,
10000 ft
,
5.8.1930
,
8♀
;
Gokča
, pr.
Babadjan-dara
,
15.5.1930
,
1♂
,
1♀
;
Abazan
, pr.
Jmerlu
,
10.8.1930
,
1♂
(all leg.
A. Schelkovnikov
);
Elenovka
,
5.6.1929
,
9♂
(leg.
Bockarev
) (all in
NHMUK
);
Aragats Mt.
,
12.6.2013
,
1♀
(leg.
A. G. Bugrov
&
I. E. Jetybayev
) (AİBÜEM);
TURKEY
: [
Kars
], Sarakancyš [Sarıkamış],
1.5.1919
,
2♂
,
1♂
nymph,
2♀
nymphs (leg.
A. Schelkovnikov
) (
NHMUK
)
;
Ardahan
,
7.5.1974
,
5♂
;
Erzurum
,
Şenkaya
,
Turnalı
,
25.6.1991
,
1♀
(leg.
E. Yıldırım
) (
EMET
)
;
Kars
,
Selim
,
Zellice Köyü
,
4.6.1980
,
4♂
,
2♀
,
1♀
nymph (AİBÜEM)
;
Erzurum
,
Horasan
,
Hızırilyas Köyü
,
1742 m
,
19.7.2011
,
1♀
(leg.
A. Hasbenli
) (
MAZM
).
Distribution.
Transcaucasia, Caucasia,
Armenia
,
Nakhichevan
,
Azerbaijan
, N.
Iran
and E.
Turkey
:
Ardahan
,
Kars
,
Erzurum
provinces (
Fig. 844
).
Remarks.
The author of this species must be Motschultsky as is the case the other species described by Motschultsky in Fischer von Waldheim, 1846 such as
Nocaracris cyanipes, Pholidoptera
pustulipes, Barbitistes foliicauda
(see Fischer von Waldheim 1846: 269, 270, 359, 360). Fischer von Waldheim gave his new species as in following format “
Nocarodes serricollis Fischer.
” in the same paper (Fischer von Waldheim 1846: 268). Subsequent researchers gave the author of
P. rubripes
as Fischer von Waldheim except
Fieber (1853: 127)
.
One male from Caucasus in the NMW was identified by Fieber as
rubripes
, but the other researchers Brunner, Ramme and Cejchan identified it as
cyanipes
. Here it is confirmed that it is
N. rubripes
and probably it is the only
type
of this species.
Motschultsky (in Fischer von Waldheim 1846: 270) in describing this species after the male gave the colors of hind tibia and tarsus as red. According to
Uvarov (1928: 150)
the inner side of hind tibia is red in male, blue-black with orange spine in female.
Bey-Bienko & Mistshenko (1951: 351)
stated that the hind tibia of this species is blueblack, black, sometimes red in male. According to
Demirsoy (1973
: 424; 1977: 81) hind tibia in both sexes is black, bluish. In fact it is quite variable, but the typical forms have red hind tibia in male and blue-black with reddish spines in the typical forms of female, as it is correctly stated by
Uvarov (1928)
(see the key above).
Avakyan (1968: 100)
synonymized
Nocaracris tibialis
Ramme
with
N. rubripes
for the first time, but it was overlooked by subsequent authors. The synonymy is here confirmed. The coloration of the hind legs of
N. tibialis
is undoubtedly within the variation of this species.
Erman & Salman’s (1990: 112) record from
Elazığ
probably belongs to another species of
Nocaracris
which should be confirmed.