The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae Author Beaver, Roger A. 161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand. Author Smith, Sarah M. 0000-0002-5173-3736 Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736 smith462@msu.edu text Zootaxa 2022 2022-08-08 5174 1 1 24 journal article 115950 10.11646/zootaxa.5174.1.1 c4bf1061-5af8-4f4a-b4ca-234b6d0ca615 1175-5326 6972960 F80F144B-D1E8-4587-A146-0BACFFE18FB6 Ips longifolia (Stebbing) Tomicus longifolia Stebbing, 1909: 26 . Ips longifolia (Stebbing) : Hagedorn 1910: 56 . Recorded from Bhutan by Schmutzenhofer (1988a) and Cognato & Sperling (2000) . Distribution. Bhutan , India ( Himachal Pradesh , Uttar Pradesh ), Pakistan . Biology. The species primarily attacks Pinus roxburghii (Pinaceae) in the subtropical conifer forest belt of the Himalayas at elevations between 500 and 2000 m ( Kirisits et al. 2002 ). Other conifers may also be attacked, but confusion with closely-related species of Ips makes most early host records unreliable. It is considered to be a secondary species, attacking severely stressed or dying trees ( Kirisits et al. 2002 ).