Order Chiroptera - Family Pteropodidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 313 350 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Cynopterus F. Cuvier 1824 Cynopterus F. Cuvier 1824 , Dentes des Mammiferes: 248 . Type Species: Pteropus marginatus E. Geoffroy 1810 Synonyms: Niadius Miller 1906 ; Pachysoma Geoffroy 1828 . Species and subspecies: 7 species with 24 subspecies: Species Cynopterus brachyotis Müller 1838 Subspecies Cynopterus brachyotis subsp. brachyotis Müller 1838 Subspecies Cynopterus brachyotis subsp. altitudinis Hill 1961 Subspecies Cynopterus brachyotis subsp. brachysoma Dobson 1871 Subspecies Cynopterus brachyotis subsp. ceylonensis Gray 1871 Subspecies Cynopterus brachyotis subsp. concolor Sody 1940 Subspecies Cynopterus brachyotis subsp. hoffeti Bourret 1944 Subspecies Cynopterus brachyotis subsp. insularum K. Andersen 1910 Subspecies Cynopterus brachyotis subsp. javanicus K. Andersen 1910 Species Cynopterus horsfieldii Gray 1843 Subspecies Cynopterus horsfieldii subsp. horsfieldii Gray 1843 Subspecies Cynopterus horsfieldii subsp. harpax Thomas and Wroughton 1909 Subspecies Cynopterus horsfieldii subsp. persimilis K. Andersen 1912 Subspecies Cynopterus horsfieldii subsp. princeps Miller 1906 Species Cynopterus luzoniensis Peters 1861 Species Cynopterus minutus Miller 1906 Species Cynopterus nusatenggara Kitchener and Maharadatunkamsi 1991 Subspecies Cynopterus nusatenggara subsp. nusatenggara Kitchener and Maharadatunkamsi 1991 Subspecies Cynopterus nusatenggara subsp. sinagai Kitchener 1996 Subspecies Cynopterus nusatenggara subsp. wetarensis Kitchener 1996 Species Cynopterus sphinx (Vahl 1797) Subspecies Cynopterus sphinx subsp. sphinx Vahl 1797 Subspecies Cynopterus sphinx subsp. angulatus Miller 1898 Subspecies Cynopterus sphinx subsp. babi Lyon 1916 Subspecies Cynopterus sphinx subsp. pagensis Miller 1906 Subspecies Cynopterus sphinx subsp. scherzeri Zelebor 1869 Subspecies Cynopterus sphinx subsp. serasani Paradiso 1971 Species Cynopterus titthaecheilus Temminck 1825 Subspecies Cynopterus titthaecheilus subsp. titthaecheilus Temminck 1825 Subspecies Cynopterus titthaecheilus subsp. major Miller 1906 Subspecies Cynopterus titthaecheilus subsp. terminus Sody 1940 Discussion: Genetic variation within the genus was discussed by Peterson and Heaney (1993) and Schmitt et al. (1995) .