Fossil Darwin wasps (Hymenoptera: Ichneumonidae) from Baltic amber Author Manukyan, Andranik R. Kaliningrad Amber Museum, Marshal Vasilevskiy Square 1, Kaliningrad 236016, Russia Author Zhindarev, Leonid A. Lomonosov Moscow State University, Leninskiye Gory, 1, GSP- 1, Moscow 119991, Russia text Palaeoentomology 2021 2021-12-14 4 6 637 647 journal article 2968 10.11646/palaeoentomology.4.6.13 5323143f-4530-4bd7-a643-9b34fc48c2d7 2624-2834 5778510 9924541E-6D36-4B6C-85B1-F12EF6830012 Rovenosa alexrasnitsyni Manukyan , sp. nov. ( Fig. 3A, B ) Holotype . GPIH No. 5046 ( ex. coll. Gröhn, no. 9186), ♀ . Well preserved complete ichneumonid wasp. Left fore leg tarsi missing; the right hind leg is separated from the body. Amber piece 26 × 21 × 7 mm . Syninclusion: Diptera , Ceratopogonidae . Etymology. This new species is named in honor of Professor Alexander Pavlovich Rasnitsyn, the prominent Russian hymenopterist and palaeontologist. Diagnosis. The new species differs from R . khalaimi in its straight ovipositor and sheaths, as well as in the by hidden of multiparous plate sensillae on the female antenna. It differs from R . rasnitsyni , which also has a straight ovipositor, by the absence of notauli, and having a propodeum with weak carina. Type locality and horizon. Baltic amber. Upper Eocene , Prussian Formation. Baltic Sea coast and amber quarry Yantarny near Kaliningrad, Kaliningrad Province , Russia . Description. Body length about 5.5 mm ( Fig. 3A ), fore wing length about 4.9 mm . Temple short, strongly narrowed behind eyes in dorsal view. Distance from lateral ocellus to eye margin is equal to ocellus diameter. Mandible and malar space are not visible. Clypeus convex, separated from face by distinct impression, without teeth on its ventral margin. Clypeus and face with fine granulated sculpture, without punctures. Occipital carina probably complete. Antennae with 28 flagellomeres, multiporous plate sensillae are hidden; flagellomeres 1–4 about 5 times as long as wide; flagellomeres 5–17 slightly elongated, about 1.2 times as long as wide; subapical flagellomeres transversе. Mesosoma moderately wide. Mesoscutum without notauli, with fine granulated sculpture; without punctures. Prepectal carina present. Sternauli and pospectal carina absent. Mesopleura punctured along mesopleural carina. Metapleural carina complete. Submetapleural carina anteriorly expanded into a broad lobe. Metapleura anteriorly finely punctuate. Propodeum with weak carina; areola about 2.5 times as long as wide ( Fig. 3B ). FIGURE 2. A , Puparium of aphidophagous syrphid in Baltic amber ( Diptera : Syrphidae : Syrphinae) (KAM No. 6361) with an emergence hole typical for Diplazontinae , ventral view. B , Right lateral view; Emergence holes of various recent parasitoids of aphidophagous syrphids ( Diptera , Syrphidae , Syrphinae). C , Diplazon pectoratorius . D , Syrphoctonus sp. E and F , Diplazon laetatorius . G , Cryptinae (gen. et sp. indet). H , Asaphes sp. (Pteromalidae) . Scale bars = 1 mm. FIGURE 3. A , Rovenosa alexrasnitsyni sp. nov. , holotype, habitus, scale bar = 1 mm. B , Mesosoma and tergites 1–2 of metasoma, left lateral view. Scale in mm. Fore wing with areolet oblique and pointed above but without stalk. Vein 2m-cu inclivous and arcuate; bulla in the half of 2m-cu directly below areolet. Pterostigma moderately broad, about 3.6 times as long as wide. Metacarp (2R1) falls short of the apex of fore wing. Vein 4Rs long, about 2.4 times as long as r-rs. Nervulus interstitial and weakly inclivous. Hind wing invisible. Hind coxae and trochanters are enlarged, length of coxae 1.1 mm , about 1.9 times as long as wide, longer than first metasomal tergite. Length of hind femur 0.7 mm , 2.5 times as long as wide. Length of hind tibia 1.5 mm , about 8 times as long as wide; spurs are short and strong. Dorsal surface of tibia without sharp spines. Claws of hind tarsi thin, weakly curved. Hind tibia and tarsi with dense, elongate and adjoined pubescence. Metasoma sharply convex behind segment 2. First metasomal tergite weakly convex dorsally in lateral view; dorsomedian carinae weak, more or less reaching post 0.7 of tergite: spiracle in 0.5 of tergite just below dorsolateral carina( Fig. 3B ). Tergites finely punctate. The body is uniformly dark brown or black, without pubescent.