Review of the Stemonosudis rothschildi species complex, with descriptions of two new species from the Indo-west Pacific Ocean (Aulopiformes: Paralepididae)
Author
Ho, Hsuan-Ching
Author
Russell, Barry
Author
Graham, Ken
Author
Psomadakis, Peter N.
text
Zootaxa
2019
2019-12-04
4702
1
216
229
journal article
24733
10.11646/zootaxa.4702.1.19
338fd90c-fb7d-4144-b0f1-49961a59b03c
1175-5326
3562801
F9EE15A8-61EB-45F6-A5A5-7660A67B91B5
Stemonosudis
Harry, 1951
Stemonosudis
Harry, 1951:32
(
Type
species:
Macroparalepis intermedius
Ege, 1933
).
Distinguishing features.
A genus of the subfamily
Paralepidinae
with body generally relatively slender; anus clearly situated before dorsal-fin origin; jaws slender; nostrils above posterior the end of maxilla (but unknown in some species because of a lack of adult specimens), and some are slightly further forward (i.e. three species treated in present study); ventral adipose fin well-developed between anus and origin of anal fin; anal-fin rays 34–50 (except
S. molesta
with 30); vertebrae more than 84 (usually more than 90).
Remarks.
Prior to the present study, total of 12 nominal species were recognized in the genus
Stemonosudis
. The two new species described herein raise the number to 14. However, based on the examination of the
type
specimens of most of species, some may be reassigned to other genera, such as
Lestidiops
.
Harry (1951
,
1953
) listed many characters in his diagnosis of
Stemonosudis
. However, most of them are commonly shared by other genera.
Stemonosudis
is similar to
Macroparalepis
Ege,
1933
in having a very slender body, but
Macroparalepis
has the anus well behind the tip of the anal fin, no ventral adipose fin and 24–28 anal-anal fin rays, whereas
Stemonosudis
has the anus just situated above posterior tip of pelvic fin, ventral adipose fin well-developed between anus and AFO, and more than 34 anal-fin rays (except for one species with 30).
Of all the other genera in the Lestidiinae,
Stemonosudis
is most similar to
Dolichosudis
Post, 1969
. Based on observations by the first author, no distinguishable character has been found between these two genera, except for the uniformly blackish color in adults of
Dolichosudis
, compared with an uneven color pattern, such as saddles, in
Stemonosudis
. These two genera are likely to be synonyms.
Stemonosudis
also shares most characters with members of
Lestidiops
Hubbs, 1916
. The two genera can be separated mainly by body shape (much more slender and narrow in
Stemonosudis
than in
Lestidiops
), shapes of jaws (more slender in
Stemonosudis
than in
Lestidiops
), and position of nostrils (above or behind posterior end of maxilla in
Stemonosudis
and well in front in
Lestidiops
). However, the three species treated here have moderately slender bodies and jaws and their nostrils are slightly in front of the end of the maxilla, conditions that are “intermediate” between these of the two genera. Further studies are needed in order to assess the validity of the two genera, and the species assigned to them may change.