The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains Author Vallejo, Luís Fernando Author Wolff, Martha text Zootaxa 2013 3722 2 101 142 journal article 10.11646/zootaxa.3722.2.1 f2c46514-f165-43d3-a574-47c55b89dcd6 1175-5326 218998 29D77A16-096D-4FC1-A5B4-9EEDF2E761A7 19. Phyllophaga ( Phyllophaga ) roscida (Burmeister, 1855) Comments. The species was originally described from a male and a female obtained in “Columbien” by Mr. Sommer. Frey (1975) stated that only the female type was found and that it is similar to P. rorulenta and to P. pastassana (Moser) from Ecuador . Revision of the non-type female deposited in MLUH showed that the mesotibial and metatibial structures are similar to the species in the subgenus Cnemarachis , however, the structure of the claws is very different to those of P. pastassana .