A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)
Author
Silva, Fernando A. B.
Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br
Author
Valois, Marcely
Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.
text
Zootaxa
2019
2019-07-08
4629
3
301
341
journal article
21174
10.11646/zootaxa.4629.3.1
9419bc9b-dbe4-426c-a34e-f0034848ff99
1175-5326
3995508
1D47B452-0A40-4BC5-9BBA-C06DEB298C9A
Scybalocanthon cyanocephalus
(
Harold, 1868
)
(
Figs. 1G
,
4G
,
5F
,
6B
)
Canthon cyanocephalus
Harold, 1868: 13
, 53 (identification key and description);
Harold 1869: 990
(catalog);
Bates 1887: 28
(taxonomic remarks);
Gillet 1911: 29
(catalog);
Schmidt 1922: 65
, 74 (diagnosis);
Balthasar 1939: 191
(redescription);
Krajcik 2006: 26
(catalog);
Krajcik 2012: 63
(catalog).
Canthon cyanocephalum
:
Blackwelder 1944: 199
(checklist);
Roze 1955: 41
(checklist).
Scybalocanthon cyanocephalus
:
Pereira & Martínez 1956: 115
, 119 (catalog and identification key);
Vulcano & Pereira 1964:
637 (catalog);
Vulcano & Pereira 1967: 554
(distribution, identification key);
Halffter & Martínez 1977: 67
(species list).
Scybalocanthon cyanocephalum
:
Vaz-de-Mello 2000: 194
(checklist).
Diagnosis
. Specimens of
S. cyanocephalus
are readily distinguishable from all other species by the pronotum with a longitudinal dark band on midline (
Fig. 1G
); elytra bicolored, with black spot on the anterior portion (
Fig. 1G
); the eighth elytral stria lacking carina on anterior portion; aedeagus with parameres elongate, subrectangular, not excavated ventrally (
Fig. 4F
); and endophallus lacking bristles or microbristles close to the FLP sclerite (
Fig. 5G
).
Description. Body.
Oval-elongate, with variable metallic sheen. Surface completely microgranulate.
Color
. Most of pronotum, disc of elytra, and ventral surface of body yellow or brown. Head, margin of pronotum and elytra, longitudinal dark band on midline of pronotum, semicircular band at the humeral portion of elytra black with green sheen.
Length
.
5.8–6.9 mm
.
Thorax
. Anterior angles of pronotum acute (approximately 85°). Lateral margin irregularly curved outward, forming an obtuse angle at the middle portion.
Elytra
. Striae thin and shiny, punctures almost inconspicuous. Eighth stria lacking carina at the anterior portion.
Aedeagus.
Parameres slightly asymmetrical (
Fig. 4G
). Dorsal margin of parameres curved inward from the basal to medial portions and curved outward at the apical portion. Ventral margin of parameres substraight, with a short denticle at the basal portion. SRP circular, with substraight handle-shaped extension (
Fig. 5F
). FLP I-shaped (
Fig. 5F
). A+SA with two superposed and elongate sclerites (
Fig. 5F
).
AS
I-shaped (
Fig. 5F
).
Type material.
Canthon cyanocephalus
Harold
,
lectotype
♂
(
here designated
;
MNHU
): “
cyanocephalus
/
Harold
/
Orinoco
” // “
Chaeridium
/
nigrocinetum
/
Mont.
/
Orinoco
/
Mont.
” // “
26363
” // “Type” // “
LECTOTYPE
♂
/
Canthon
/
cyanocephalus
/
Harold
/ des.
F. Z. Vaz-de-Mello
”.
Paralectotype
♂
(
MNHU
): “
cyanoce
/
phalus Har.
”// “
26363
” // “Type” // “
PARALECTOTYPE
/
Canthon
/
cyanocephalus
/
Harold
/ des.
F. Z. Vaz-de-Mello
”
.
Non-type material [
22 males
,
22 females
]:
BRAZIL
:
AMAPÁ
: Amapari, Rio Mariru (
51°57′13″W
,
0°46′10″N
), N. Degallier—
1♂
(
CEMT
). PARÁ: Monte Dourado, rainforest (52°67’W, 00°85’S),
iv.2006
, pitfall dung, T.A. Gardner & M.I.M. Hernández—
1♂
(
CEMT
); Monte Dourado,
Eucaliptus
forest (
52°51’W
,
0°57’S
),
19.v.
2008
, 180 m, E. Bordoni—
2♂
2♀
(
CEMT
).
FRENCH GUIANA
: Cayenne (
4°54’3’’N
,
52°18’12’’W
),
x.1918
, [anonymous]—
1♂
(
CMNC
).
TRINIDAD AND TOBAGO
: COUVA-TABAQUITE-TALPARO, Quesnell Farm,
13 km
S
Arima
,
2 km
N
Talparo
,
50 m
(
10°31’48’’N
,
61°16’26’’W
),
12–22.vi.1993
, S. & J. Peck, rainforest, flight interception trap (1993-28)—
1♂
1♀
(
CMNC
); Quesnell Farm,
13 km
S
Arima
,
2 km
N
Talparo
,
50 m
(
10°31’48’’N
,
61°16’26’’W
),
22.vi–8.vii.1993
, S. & J. Peck, rainforest, flight interception trap (1993-46)—
1♀
(
CMNC
); EAST- ERN
TOBAGO
, Gilpin Trace,
10 km
NE Roxborough,
400–500 m
(
11°17’N
,
60°36’W
),
26–30.vi.1993
, S. & J. Peck, flight interception trap (1993-50)—
1♂
3♀
(
CMNC
); SAN JUAN-LAVANTILLE, above Loango Village, Maracas Valley,
600 m
(
10°44’1’’N
,
61°26’10’’W
),
9–22.vi.1993
, S. & J. Peck, mountain rainforest, flight interception trap (1993-22)—
1♀
(
CMNC
); above Loango Village, Maracas Valley,
600 m
(
10°44’1’’N
,
61°26’10’’W
),
22.vi–6.vii.1993
, S. & J. Peck, mountain rainforest, flight interception trap (1993-43)—
1♂
(
CMNC
); Maracas Bay (
10°45’13’’N
,
61°25’58’’W
),
23–30.vi.1976
, J.D. Glaser—
1♂
(
CMNC
); Simla Research Station,
8 km
N
Arima
,
240 m
(
10°42’20’’N
,
61°17’40’’W
),
6–10.vi.1993
, S. & J. Peck, tropical forest, flight interception trap (1993- 11)—
1♂
(
CMNC
); TUNAPUNA-PIARCO, Andrews Trace,
16 km
N
Arima
,
620 m
(
10°43’45’’N
,
61°18’40’’W
),
7–24.vi.1993
, S. & J. Peck, upper mountain rainforest, flight interception trap (1993-14)—
2♂
1♀
(
CMNC
); Blan- chisseuse Road,
610 m
(
10°43’N
,
61°18’W
),
14.ii.1987
, M.S. Adams—
1♀
(
CMNC
); Lalaja Trace,
19 km
N
Arima
,
650 m
(
10°44’26’’N
,
61°18’41’’W
),
8–24.vi.1993
, S. & J. Peck, mountain rainforest, flight interception trap (1993- 20)—
2♂
2♀
(
CMNC
); Lalaja Trace,
19 km
N
Arima
,
650 m
(
10°44’26’’N
,
61°18’41’’W
),
24.vi–7.vii.1993
, S. & J. Peck, mountain rainforest, flight interception trap (1993-59)—
1♂
(
CMNC
); Morne Bleu,
823 m
(
10°43’N
,
61°17’W
),
4.viii.1969
, H. & A. Howden—
1♀
(
CMNC
); Mount Saint Benedict,
Tunapuna
,
550 m
(
10°40’51’’N
,
61°23’36’’W
),
5–21.vi.1993
, S. & J. Peck, summit rainforest, flight interception trap (1993-06)—
1♂
2♀
(
CMNC
); Mount Saint Benedict,
Tunapuna
,
550 m
(
10°40’51’’N
,
61°23’36’’W
),
5.vi.1993
, S. Peck, treebase litter and seeds, tree base litter (1993-07)—
1♂
(
CMNC
); Simla Research Station,
8 km
N
Arima
,
260 m
(
10°42’20’’N
,
61°17’40’’W
),
6–14.vi.1993
, S. & J. Peck, tropical forest, flight interception trap (1993-09)—
1♂
(
CMNC
); Simla Research Sta- tion,
Arima
Valley, North Range,
244 m
(
10°41’34’’N
,
61°17’22’’W
),
24–29.v.2000
, A.S. Ramsdale, premontane rainforest, pitfall trap, dung—
1♀
(
CMNC
); Simla,
5 mi.
N
Arima
(
10°43’7’’N
,
61°17’58’’W
),
18.viii.1969
, H. & A. Howden—
1♂
1♀
(
CMNC
); Simla,
5 mi.
N
Arima
(
10°43’7’’N
,
61°17’58’’W
),
20.viii.1969
, H. & A. Howden—
1♂
(
CMNC
).
VENEZUELA
:
BOLÍVAR
,
26 km
N Guasipati (
7°41’35’’N
,
61°57’19’’W
),
24.vi–12.vii.1987
, S. & J. Peck, sandy seasonally humid forest, flight interception trap—
1♂
4♀
(
CMNC
); Anacoco (
6°43’0’’N
,
61°8’00’’ S
),
6.viii.2006
, Curso NM 2006–
1♀
(
CEMT
); Anacoco (
6°43’0’’N
,
61°8’00’’ S
),
6.viii.2006
, NM Curso 2007–
1♂
(
CEMT
).
Distribution.
Known from
Brazil
(
Amapá
, Pará),
French Guiana
,
Trinidad and Tobago
and
Venezuela
(
Fig. 6B
).
Endemism areas
:
Brazilian sub-region
: Boreal Brazilian dominion: Guianan Lowlands and Roraima provinces; Mesoamerican dominion:
Trinidad
province (see
Morrone 2014
; fig. 12).