Nearctic species related to Diogmites angustipennis Loew (Diptera: Asilidae) Author Barnes, Jeffrey K. text Zootaxa 2010 2545 1 22 journal article 10.5281/zenodo.196750 a17da0e2-b742-4c18-b255-8655d9893934 1175-5326 196750 Diogmites bilobatus sp. nov. ( Fig. 2 ) Diagnosis. Male 16.2–25.3 mm long (mean = 21.1 mm); female 17.0–26.5 mm long (mean = 22.5 mm). Average antennomere:scape ratios 1.0:1.6:4.1. Male wing 11.7–16.5 mm long (mean = 14.3 mm); female wing 12.2–18.4 mm long (mean = 15.6 mm). Phallus bilobed ( Fig. 2 a). Female abdominal tergites 1–6 pollinose and opaque dorsally; tergites 7–8 glabrous dorsally, opaque laterally ( Fig. 2 b). Description. Male body, excluding antennae, 16.2–25.3 mm long (mean = 21.1 mm); female 17.0–26.5 mm long (mean = 22.5 mm); golden brown to black in ground color; golden tomentose almost throughout. Head width 1.7 times eye height. Face golden tomentose. Mystax a dense row of long, yellowish-white setae on extreme lower margin of face; upper face lacking setae. Frons golden tomentose, with scattered thin, pale, hair-like setae. Vertex brown, with scattered, thin, pale to black, hair-like setae. Ocellar triangle tuberculate, with two long, strong, black setae and several short, thin, pale to black, hair-like setae. Postocular setae long, strong, pale. Postgena and stipes with dense vestiture of long, thin, pale, hair-like setae. Proboscis dark brown, somewhat laterally compressed, more than five times as long as deep; deepest near base, tapered to rounded apex. Palpus about as long as scape and pedicel combined, brown, bearing many long, pale, hairlike setae. Antenna brown, about 0.43 times head width; pedicel with several black setae dorsally and ventrally; flagellum with several black setae dorsally on basal half; antennomere:scape ratios 1.0:1.6:4.1. FIGURE 2. Diogmites bilobatus sp. nov. a, phallus, dorsal view. b, ovipositor, lateral view. c, United States distribution, by county or parish. Thorax with cervical sclerites covered with vestiture of long, pale setae. Antepronotum with row of long, slender, pale setae along anterior margin; postprontoum bearing sparse, long, pale, hair-like setae laterally; postpronotal lobe bearing 1–2 strong, black or pale setae and several short pale, hair-like setae. Proepisternum covered with short, thin, pale, hair-like setae. Prosternum lacking setae. Scutum golden brown to dark brown in ground color, golden tomentose laterally, with three dorsal longitudinal, golden tomentose vittae, lacking distinct dark makings; mid-dorsal vittae very narrow; anterior third and golden tomentose areas covered with short, black setae; with two strong, black notopleural setae, two supraalar setae, two postalar setae, 2–4 strong dorsocentral setae, and two strong, black setae on postalar callus. Scutellum low, not convex, with two strong, black marginal setae. Mediotergite and anatergite bare; katatergite with fan of 6–7 long, slender, pale setae. Mesopleural sclerites lacking setae. Legs yellowish brown. Coxae golden tomentose; fore and mid coxae with many long, slender, pale anterior setae; hind coxa with 1–2 strong, pale lateral setae and several thin, hair-like anterior and lateral setae. Trochanters golden tomentose; fore and mid trochanters with ventrolateral dark spots; hind trochanters with ventrolateral dark line and many, short, stout, pale, ventral setae. Femora, tibiae, and tarsi covered with many, short, thin, black setae. Fore femur with single, strong, stout, black anterior seta near basal fifth; row of 3–5 similar stout setae prosteriorly; and irregular row of shorter, stout, black setae anteroventrally. Mid femur with single strong, stout, black, anterior seta near basal fifth and row of shorter black anterior setae; single strong, stout, black posterior seta near apex and row of 4–5 shorter black posterior setae; and single strong, stout, black ventral seta near base. Hind femur with row of 4–8 short, stout, black anterior setae; row of 2–5 short, stout, black anteroventral setae; and similar row of posteroventral setae. All tibiae with several long, black, preapical setae. Fore tibia with anterodorsal, posterodorsal, and posteroventral rows of 5–8 short, stout, black setae; single short, stout, black ventral seta about one third distance from base; and single long, strong, black ventral seta about one third distance from apex. Mid tibia with dorsal, anterodorsal, anteroventral, and posteroventral rows of short, stout, black setae, only posteroventral row extending from base to apex, other rows terminating about halfway from base; with two long black anterodorsal and two long, black ventral setae. Hind tibia with anterodorsal, posterodorsal, anteroventral, and posteroventral rows of long, black setae; posterodorsal and distal anteroventral setae longest; posteroventral row extending only about halfway from base of tibia. Tarsi darkened apically, with many strong, black setae; first tarsomere about equal in length to following three tarsomeres. Fore tarsus about equal in length to fore tibia; mid and hind tarsi shorter than mid and hind tibiae, respectively. Claws black; pulvilli pale yellowish-brown. Wing 11.7–16.5 mm long in male (mean = 14.3 mm); 12.2–18.4 mm long in female (mean = 15.6 mm). Membrane lightly infuscate. Halter with stem and knob golden brown, knob somewhat darker. Abdomen golden brown to dark brown or black in ground color. Tergites 2–5 with diffuse, diagonal, dark brown to black patches laterally, subtended posteriorly by silver or golden tomentose patches. Phallus bilobed ( Fig. 2 a). Female abdominal tergites 1–6 pollinose and opaque dorsally; tergites 7–8 glabrous dorsally, opaque laterally ( Fig. 2 b). Both sexes with all abdominal sternites pollinose and opaque. Discussion. Diogmites bilobatus sp. nov. is much more consistent in size than is D. angustipennis , and on average it is slightly shorter in length, but its range of color variation is more extreme. Some specimens, especially those from southern Texas, have nearly black abdominal tergites. Specimens collected at more northerly sites tend to be lighter in color. Abdominal tergites of specimens collected in Doña Ana County in southwestern New Mexico range across the entire spectrum, from golden orange to nearly black. The known range of D. bilobatus sp. nov. extends from northern Kansas south through Oklahoma and Texas to northeastern Mexico , and from Louisiana and southeastern Arkansas west to Arizona and southern California ( Fig. 2 c). Labels on museum specimens indicate that adults are active from late May to mid October, but most museum specimens bear labels dated July, August, or September. The earliest and latest records refer to specimens collected in far southern reaches of the species’ range. G. S. Forbes (in litt.) collected many specimens in New Mexico and made the following observations: The new species probably originally inhabited the Rio Grande floodplain before they channeled the river and tore out the bosques. At the river it now seems to occur in flat sandy areas between the levee roads and the river edge. Although the sites are periodically mown and otherwise "managed" by the International Boundary and Water Commission, they often get quite weedy, with Astragalus , small tamarisks, Salsola , Bermuda grass, and rabbit brush, with Salix exigua along the bank. The Diogmites is a rather low flier, perching on weeds, grass stems, and often on the Salix . It is somewhat wary and doesn't perch for long at any one spot. Other perching sites I noted were Chrysothamnus and mesquite. The species has also extended into agricultural situations and was frequently found in alfalfa fields, where there were doubtless many prey items. The fly is also present in residential areas. It often perched on Bermuda grass. The only record I have from outside the Mesilla Valley was a male from the south end of the USDA Jornada Experimental Range, on the HQ road 6.6 mi. NW of Rt. 70, 19 vii 1982. This is an area of mixed creosotebush-snakeweed-yucca scrub with saltbush and mesquite along the washes. Along the road is a grassy area with tobosa grass and Scleropogon brevifolia . The species was locally common at times but I see it wasn't recorded at several of the riparian sites I collected, nor every time during the flight season. The most abundant I found it was on 2 Aug. 1982 on the E levee rd. 2 mi . (NW) of Picaho Ave, where I recorded 12 in about 1 1/4 hours. The only prey record I have is a female Efferia albibarbis . Triorla interrupta , a Machimus nr. erythocnemius , Megaphorus pulchrus , and Efferia albibarbis were found with it in weedy areas, and Proctacanthella cacopiloga , Efferia luna , Scleropogon duncani , Proctacanthus nearno , and Promachus giganteus occurred in some of the same sites at the proper time, although I didn't necessarily record them with the Diogmites . Some museum specimens examined during the course of this study have the following prey items pinned beneath them: Araneae : Lycosidae , Pirata sp, Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Orthoptera : Acrididae , Syrbula admirabilis (Uhler) , Texas, Live Oak County, Choke Canyon State Park, 4.X.1997 (BYU). Tettigoniidae (nymph), Texas, Brazos County, College Station, 12.VIII.1932 (USNM). Hemiptera : Coreidae , Anasa andresii (Gurin-Mneville) , Texas, Hidalgo County, 8.IX.1933 (USNM). Coreidae , Chariesterus antennator Fabricius , Texas, Brazos County, College Station, 20.VIII.1933 (TAMU). Coleoptera : Scarabaeidae , Onthophagus sp., Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Histeridae , Texas, Brazos County, College Station, 10.VIII.1932 (USNM). Diptera : Asilidae , Efferia sp., Texas, Brazos County, College Station, 8.VIII.1932 (USNM). Asilidae , Proctacanthella cacopiloga (Hine) , Texas, Brazos County, College Station, 19.VIII.1932 (USNM). Syrphidae , Vollucella mexicana Macquart , Texas, Cameron County, 8.VIII.1933 (USNM). Syrphidae , Eristalis sp., Texas, Hidalgo County, Weslaco, 7.VIII.1933 (USNM). Calliphoridae , Lucilia illustris (Meigen) , Texas, Hidalgo County, Weslaco, 12.VIII.1933 . Tachinidae , Texas, Madison County, 22.VIII.1932 (USNM). Hymenoptera : Anthophoridae , Melissodes sp., Texas, Brazos County, College Station, 3.VIII.1932 (USNM). Hymenoptera , Anthophoridae , Melissodes sp., Texas, Hidalgo County, Donna, 8.X.1933 (USNM). Apidae , Apis mellifera Linnaeus , Texas, Bexar County, 2.X.1946 (USNM). Apidae , Apis mellifera Linnaeus , Texas, Bexar County, 6.VI.1934 (TAMU). Apidae , Apis mellifera Linnaeus , Texas, Bexar County, 28.X.1946 (USNM). Apidae , Apis mellifera Linnaeus , Texas, Brazos County, College Station, 14.X.1945 (USNM). Apidae , Apis mellifera Linnaeus , Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Halictidae , Halictus sp., Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Ichneumonidae , Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Ichneumonidae , Texas, Live Oak County, Choke Canyon State Park, 4.X.1997 (BYU). Nyssonidae, Gorytinae , Texas, Hidalgo County, Weslaco, 7.VIII.1933 (USNM). Pompilidae , Texas, Brazos County, College Station, 30.VII.1932 (USNM). Sphecidae , Podium sp., Texas, Hidalgo County, Weslaco, 12.VIII.1933 (USNM). Sphecidae , Tachytes sp., Texas, Brazos County, College Station, 15.VII.1932 (USNM). Tiphiidae , Texas, Brazos County, College Station, 17.VIII.1932 (USNM). Scoliidae , Campsomeris tolteca Saussure , Texas, Hidalgo County, 12.VII.1933 (USNM). Vespidae , Polistes , Texas, Brazos County, College Station, 8.VIII.1932 (USNM). Lepidoptera : Libytheidae , Libytheana carinenta (Cramer) , Texas, Cameron County, 8.VIII.1933 (USNM). One Blythe, California, specimen in the Charles H. Martin collection at the Florida State Collection of Arthropods bears a label stating, “This species common Blythe area. They kill thousands of honey bees. It was captured in a cotton field on 22 July 1949 . Type material. Holotype : The male holotype is deposited in the CAS collection. It is labeled ‘San Antonio, Tex’, VI-25-42 ’, E S Ross’; ‘ Diogmites , symmachus, Lw – ’76, det. J. Wilcox’, ‘HOLOTYPE’, ‘ DIPTERA : ASILIDAE’, ‘ Diogmites bilobatus’, ‘Barnes’. It measures 20.4 mm long and wings are 13.1 mm long. The antennomere:scape ratios are 1.0:1.7:3.8. Paratypes : UNITED STATES . ARIZONA: Cochise County, 8 mi . NE of Douglas, 4.VIII.1940 , ɗ ( UCR ); 8 mi . E of Douglas, 8.VIII.1958 , ɗ ( UCDC ); 27 mi . E of Douglas, 9.VIII.1966 , ɗ ( UCR ); Miller Canyon, Huachuca Mountains, 5000', 9.VII.1974 , 2 Ψ ( CUIC ); 26.VII.1974 , Ψ ( CUIC ); San Bernardino Ranch, 3750', August, Ψ ( CAS ). Coconino County, Flagstaff, 25.VII.1950 , 2 Ψ ( AMNH ). Maricopa County, Mesa, VI.1931 , ɗ, Ψ ( BYU ); 17.VII.1950 , Ψ ( EMUS ). Mohave County, Mohave Valley, 1.VI.1980 , ɗ, Ψ ( UAIC ); 15.VII.1980 , 2 ɗ ( UAIC ); 25.VII.1980 , ɗ ( UAIC ). Pima County, 20.IX.1925 , Ψ ( CAS ); Tucson, 20.X.1927 , Ψ ( CAS ); X.1931 , Ψ ( CAS ); 21.IX.1938 , Ψ ( CAS ); 4.VIII.1953 , ɗ ( UAIC ); 13.VIII.1954 , Ψ ( UCDC ); 29.VIII.1954 , Ψ ( UAIC ); 16.VIII.1955 , Ψ ( UAIC ); laying eggs on soil, 31.VII.1966 , Ψ ( UAIC ). Yuma County, Roll, 19.VII.1954 , Ψ ( UAIC ). ARKANSAS: Pike County, Daisy, 4.VII.1974 , ɗ (WTAM). CALIFORNIA: Riverside County, Blythe, cotton field, killing honey bees, 22.VII.1949 , ɗ (FSCA). COLORADO: county unknown, Walker Hill, 6.VIII.1933 , 2 ɗ (CSUC, WSU). Alamosa County, Alamosa, Summer 1964, ɗ (FSCA). Baca County, S of Lamport, 2.VIII.1933 , ɗ, Ψ (CSUC). Bent County, Las Animas, 2.VIII.1921 , Ψ (CSUC); 12.IX.1933 , Ψ (WSU); 3887', 6.IX.1938 , Ψ (SEMC); Rd CC, SW of Caddoa, 17.VIII.1995 , Ψ (CSUC); Rd 26, along Caddoa Cr., 17.VIII.1995 , ɗ (CSUC). Crowley County, Ordway, 19.VII , Ψ (CSUC); 6.VIII.1933 , 2 Ψ (CSUC); 11.VIII.1934 , Ψ (CSUC); 12.VIII.1934 , Ψ (CSUC); 23.VIII.1936 , ɗ (CSUC); 6.VIII.1937 , ɗ (WSU). Las Animas County, Purgatoire River, Bent Canyon, 2.VII.1989 , ɗ (CSUC). Otero County, CO Rt. 109, Purgatoire R., 11.VIII.1990 , 2 ɗ, Ψ (CSUC). Prowers County, Hartman, 23.VIII.1961 , ɗ, Ψ (KSUC); Lamar, 25.VIII.1925 , 2 ɗ, 3 Ψ (UMMZ); Lamar, Arkansas River, 15.IX.1992 , Ψ (CSUC); Lamar, Willow Creek Park, 15–16.VIII.1998 , Ψ (CSUC). Pueblo County, Boone, 5000', 17.VIII , Ψ (KSUC). KANSAS: Hamilton County, 4000', 6.VIII.1928 , ɗ (KSUC); 8 km WNW of Syracuse, 25.VII.1991 , Ψ (CMNH). Meade County, 14.VIII.1945 , 2 ɗ, Ψ (SEMC). Morton County, 2800', 5.VIII.1911 , Ψ (OSEC). Rooks County, 1775 ft ., 9.VIII.1912 , ɗ (OSU). Seward County, 2600', 18.VIII.1911 , 6 ɗ, 3 Ψ (OSU). Wallace County, 19.VIII , ɗ (KSUC). LOUISIANA: Acadia Parish, Crowley, 22.IX.1911 , Ψ (USNM). Calcasieu Parish, Sulphur, 23.VIII.1992 , Ψ (BYU); 14 mi . S Vinton, 8.IX.1916 (USNM). Cameron Parish, Cameron , 2.VII.1905 , ɗ (OSU); 10.VII.1905 , 3 Ψ (OSU); Martin Beach off Hwy 82, 13 .VIII.2003, Ψ (CSUC). Lafayette Parish, Lafayette, Ψ (USNM); 11.X.1932 , Ψ (USNM); Southwestern Lafayette, 8.X.1938 , ɗ (USNM); 5.XII.1938 , ɗ (USNM). Madison Parish, Tallulah, Ψ (OSEC). NEW MEXICO : county unknown, northern New Mexico , ɗ (USNM). Bernalillo County, Albuqueque, VIII.1894 , ɗ, Ψ (OSU). Chaves County, 8.VII.1927 , ɗ (SEMC); Chaves, 6.VIII , ɗ (OSU). Doña Ana County, alfalfa, 3.VIII.1956 , Ψ (NMSE); alfalfa, 16.VIII.1956 , Ψ (NMSE); swept from alfalfa, 9.VIII.1956 , ɗ (NMSE); 5 mi . N of Anthony, 16.VII.1958 , ɗ (WFBM); Las Cruces, 23.VIII , ɗ (USNM); 28.VIII.1937 , Ψ (USNM); 31.VII.1961 , Ψ (NMSE); Las Cruces, Mesquite Drive, 6.X.1975 , Ψ (NMSB); Las Cruces, Rio Grande, E levee road, 2 mi . N of Picacho, 25.VII.1982 , Ψ (NMSB); 29.VII.1982 , 8 ɗ, 3 Ψ (BYU, NMSB); 2.VIII.1982 , 9 ɗ, Ψ (EMUS, NMSB); 23.VIII.1982 , 2 Ψ (EMUS, NMSB); Las Cruces, Rio Grande, W levee road, 1.5 mi. of Picacho Ave, 6.IX.1982 , Ψ (NMSB); Rio Grande, 1 mi . E of San Miguel, 3900', 18.VIII.1993 , Ψ (NMSB); Las Cruces, yard, 2 mi . N of NMSU, 12–14. VII.1982, 2 Ψ (NMSB); 13.VII.1982 , ɗ (NMSB); 18.VII.1982 , ɗ (NMSB); 2 mi . N of NMSU campus, 22.VII.1982 , ɗ (BYU); 2 mi . N of NMSU campus, 3900', 25.VII.1982 , ɗ (NMSB); 2 mi . N NMSU campus, 24.VII.1983 , ɗ (NMSB); 10 mi . S NMSU Exp. Farm, 6.VIII.1982 , ɗ, 4 Ψ (NMSB); 10 mi . S NMSU Exp. Farm, 3900', 7.VIII.1982 , Ψ (NMSB); playa 6.6 mi. N Rt. 70/82, 0.5 mi. E Joranda Rd., 4200', 19.VII.1982 , ɗ (NMSB); Masilla Dam, 3800', 9.VIII.1979 , Ψ (NMSB); 16.VII.1982 , Ψ (NMSB); 31.VIII.1982 , ɗ (NMSB); Mesilla Park, alfalfa, 17.VIII.1962 , Ψ (NMSB); White Sands, 7.VIII.1931 , ɗ (KSUC). Eddy County, Carlsbad, 5.VIII.1931 , ɗ (KSUC); 2.IX.1967 , ɗ, 2 Ψ (UCR); Loving, 19.VI.1963 , ɗ (NMSE); Malaga, 11.VII.1936 , ɗ (CAS). Roosevelt County, 2.VII.1968 , ɗ (UCR); Dora, 7.VIII.1965 , Ψ (CAS). San Miguel County, Pecos, 16.VIII.1938 , Ψ (AMNH). Sierra County, Percha Dam State Park, Rio Grande River, 17.VII.1989 , ɗ (CSUC). Socorro County, La Joya Wilderness, 31.VII.1993 , ɗ (CSUC). OKLAHOMA: Beckham County, US 66 ( I-40 ) 10 Mi. E of Erick, 10.VIII.1969 , Ψ (CUIC). Logan County, near Coyle, banks of the Cimarron R., 18.VIII.1959 , 3 ɗ, 4 Ψ (OSEC). McCurtain County, Sherwood, 27.VI.1937 , Ψ (OSEC). TEXAS: county unknown, Buff Lake, 23.VIII.1968 , Ψ (ESUW). Aransas County, 6.VIII.1928 , 3 ɗ, 2 Ψ (SEMC, UAAM); 6.VIII.1928 , 2 ɗ (UAAM); Goose Island State Park, 11.VIII.1985 , ɗ, Ψ (TAMU). Baylor County, Lake Kemp, below Dam, 20.VIII.1969 , ɗ (ESUW). Bexar County, ɗ, Ψ (CAS); 2.X.1932 , ɗ, Ψ (CAS, FSCA); 2–8. X.1932, 3 Ψ (CAS); 6.VI.1934 , ɗ (TAMU); 2.X.1946 , Ψ (USNM); 28.X.1946 , Ψ (USNM); 1.VII.1952 , ɗ (EMEC); San Antonio, 22–31.VIII , Ψ (CMNH); 25.VI.1938 , 2 Ψ (SEMC, UAAM); 25.VI.1942 , ɗ (CAS); 20 mi . N of San Antonia, 22.VI.1942 , Ψ (CAS). Blanco County, Blanco, 22.VII.1962 , Ψ (TAMU); Pedernales Falls State Park, 28.IX.1991 , Ψ (BYU). Bosque County, Valley Mills, 24.VII.1932 , Ψ (USNM). Brazoria County, Alvin, 9.XI.1904 , ɗ (AMNH). Brazos County, College Station, 13.VIII.1921 , Ψ (EMEC); 5.VII.1932 , Ψ (USNM); 15.VII.1932 , ɗ (USNM); 21–22.VII.1932 , ɗ, 2 Ψ (USNM); 28.VII.1932 , ɗ, 2 Ψ (USNM); 30.VII.1932 , ɗ (USNM); 3.VIII.1932 , ɗ (USNM); 8.VIII.1932 , 4 ɗ (USNM); 11.VIII.1932 , ɗ, 2 Ψ (USNM); 12.VIII.1932 , 2 ɗ (USNM); 15.VIII.1932 , Ψ (USNM); 17.VIII.1932 , 2 ɗ, 2 Ψ (USNM); 2– 18. VIII.1932, 8 Ψ (USNM); 18.VIII.1932 , ɗ (TAMU); 18–19. VIII.1932, 3 ɗ, 2 Ψ (USNM); 21.VIII.1932 , ɗ (USNM); 20.VIII.1933 , 2 ɗ (CAS, TAMU); 26.VII.1933 , Ψ (USNM); 14.X.1945 , Ψ (USNM). Brewster County, Big Bend National Park, Rio Grande Village, trailer camp, 14.VIII.1970 , Ψ (FSCA). Cameron County, 3.VII.1933 , 2 ɗ, Ψ (USNM); 8.VIII.1933 , 5 ɗ (USNM); 18.VIII.1933 , ɗ (USNM); Brownsville, June, Ψ (SEMC); 23.X.1960 , ɗ (FSCA); 10.VII.1970 , Ψ (TAMU); 27.V.1989 , ɗ (UCR); 8 mi . E of Brownsville, 18.VII.1962 , Ψ (KSUC); 20 mi . E of Brownsville, alkaline flats, 30.VI.1968 , Ψ (FSCA); Harlingen, 4.IX.1945 , ɗ, 2 Ψ (ISUI); 3.X.1945 , Ψ (ISUI); 8.X.1945 , 2 ɗ (ISUI); Laguna Madre, 25 mi . SE Harlingen, 26.IX.1945 , 5 ɗ, 3 Ψ (ISUI); 9.X.1945 , 2 ɗ, 2 Ψ (ISUI). Childress County, Lake Childress, 24.VIII.1948 , ɗ, Ψ (AMNH). Collingsworth County, Salt Fork, Red River, Hwy 83, 11 .VIII.2003, ɗ, 2 Ψ (CSUC). Culberson County, 5 mi . E of Van Horn, 12.IX.1970 , Ψ (UCDC). Dallas County, 13.VII.1940 , Ψ (WIRC); Dallas, 22.VIII.1905 , Ψ (USNM); 14.IX.1918 , Ψ (UMMZ); 20.IX.1918 , 2 Ψ (UMMZ). Dickens County, N of Wichita River, 6 mi . SW Dumont, 9 July , Ψ (ESUW); Spur, 11.IX.1931 , Ψ (USNM). Dimmit County, Carrizo Springs, 19.VIII.1994 , Ψ (BYU). El Paso County, El Paso, 29.VII.1934 , 2 ɗ, 2 Ψ (CAS); Tornillo, mesquite area, 15.VII.1948 , Ψ (UCDC); Ysleta, 4.IX.1927 , Ψ (CUIC); 20.VII.1955 , ɗ (WSU); 3.5 mi. S Ysleta, 3800', 19.VIII.1952 , Ψ (EMEC). Gaines County, Seminole, 4.IX.1990 , Ψ (ESUW). Galveston County, Dickinson, VI.1929 , Ψ (KSUC); Galveston Island, 21.VII.1962 , 3 ɗ (KSUC); Port Bolivar, 22.VII.1962 , ɗ (KSUC). Garza County, Justiceburg, Double Mt. Fork, Brazos R. & Hwy 84, 6 .VIII.1993, Ψ (BYU). Gillespie County, Fredericksburg, 1962, Ψ (ISUI). Gray County, 20.IX.1969 , ɗ (ESUW); 3.X.1969 , ɗ (ESUW). Grayson County, 2.VIII.1969 , Ψ (UCR). Harris County, Bellaire, 11.VIII.1957 , ɗ (EMEC); Houston, 1.IX.1945 , Ψ (EMEC); 6.X.1990 , Ψ (BYU); Lake Houston, 13.VII.1978 , Ψ (FSCA). Haskell County, Lake Stamford, 29.VIII.1968 , ɗ (ESUW). Hidalgo County, 9.VII.1932 , ɗ (USNM); 26–29. VI.1933, 2 ɗ (USNM); 12.VII.1933 , Ψ (USNM); 1.VIII.1933 , Ψ (USNM); 23.VIII.1933 , Ψ (USNM); 8.IX.1933 , Ψ (USNM); Bentsen, Rio Grande Pk., 9.VI.1981 , ɗ, Ψ (CSUC); Donna, 8.X.1932 , ɗ (USNM); 22.VIII.1933 , ɗ (USNM); 24.IX.1933 , ɗ (TAMU); 31.IX.1933 , ɗ (OSU); Edinburg, FM 2812 & Doolittle Rd., 12.X.1991 , ɗ (BYU); McAllen, 11.IX.1965 , ɗ (ISUI); 19.IX.1969 , Ψ (ISUI); 14.X.1969 , ɗ (ISUI); McAllen, valley bot. garden, 27.VIII–8.IX.1973 , Ψ (FSCA); Pharr, 20.IX.1947 , ɗ, Ψ (ISUI); Weslaco, 2.VII.1933 , Ψ (USNM); 4.VII.1933 , ɗ, Ψ (USNM); 27.VII.1933 , 2 ɗ (USNM); 3.VIII.1933 , ɗ (USNM); 3–7. VIII.1933, 2 Ψ (USNM); 7.VIII.1933 , 5 ɗ, 2 Ψ (USNM); 12.VIII.1933 , 5 ɗ, 6 Ψ (USNM); 14.VIII.1933 , 2 ɗ (USNM). Hockley County, Levelland, 7.IX.1969 , Ψ (ESUW). Jeff Davis County, Ft. Davis, Limpia Creek Campground, 28.VII.1967 , ɗ, Ψ (CUIC). Jefferson County, Beaumont, 10.VIII.1917 , Ψ (CUIC). Jim Wells County, Nueces River at Texas 359, 17 .VI.1966, Ψ (FSCA). Kleberg County, Kingsville, 6.VII.1921 , Ψ (KSUC); 11.VII.1950 , ɗ (CAS); Buffin Bay, Riviera Beach, 17.VI.1948 , Ψ (INHS). Knox County, 9.VIII.1969 , ɗ (ESUW). Lavaca County, Hallettsville, 28.IX.1953 , Ψ (CAS). Lee County, Old Dime Box, 19.VI.1956 , ɗ (CUIC). Live Oak County, Choke Canyon State Park, Frio River spillway, 4.X.1997 , 4 Ψ (BYU); Choke Canyon State Park, 4 mi . W of Three Rivers, South Shore Campground, 19.X.1991 , Ψ (BYU). Lubbock County, 16.IX.1969 , Ψ (ESUW); 28.IX.1969 , 2 Ψ (ESUW); Lubbock, 2.VIII.1962 , ɗ (ESUW); 15.IX.1967 , ɗ (ESUW); 17.IX.1967 , ɗ (ESUW); 27.IX.1967 , ɗ (ESUW); 4.IX.1969 , Ψ (ESUW); 7.IX.1969 , Ψ (ESUW); 16.X.1969 , Ψ (ESUW). Madison County, 2.VIII.1930 , ɗ (TAMU); 22.VIII.1932 , 4 ɗ, 2 Ψ (OSU, USNM); X.1932, 2 ɗ, 2 Ψ (CAS, TAMU, USNM); 14.VIII.1933 , Ψ (USNM). Milam County, Thorndale, 2.X.1950 , Ψ (AMNH). Montgomery County, Montgomery, 18–21.VIII.1975 , Ψ (UCDC). Motley County, 12.VIII.1969 , 2 ɗ (ESUW, MEMU). Pecos County, Girvin, 8 mi . NW of Hwy 11, 28–30 .VIII.1992, Ψ (BYU); Hwy 10 & Pecos River, 21.IX.1984 , Ψ (TAMU). Randall County, Canyon, 13.VIII.1969 , ɗ (KSUC); Palo Duro Canyon State Park, 22.VIII.1990 , 6 ɗ, 2 Ψ (BYU); Palo Duro Canyon State Park, Hackberry Campground, 20.IX.1999 , Ψ (BYU). Reeves County, Balmorhea, 16.VIII.1982 , ɗ (TAMU). San Patricio County, Mathis, 22.VIII.1932 , Ψ (CAS); R. & B. Welder Refuge, 26.VIII.1979 , Ψ (BYU). Schleicher County, 17.1 mi. W of Eldorado on I90, 14 .VIII.1982, Ψ (TAMU). Starr County, 13.VIII.1937 , Ψ (USNM). Tarrant County, 14.VII.1968 , Ψ (ESUW). Terry County, 27.IX.1969 , Ψ (ESUW). Travis County, Austin, 6.X.1901 , Ψ (USNM); 25.IX.1965 , Ψ (U C D C). Va l Ve r de County, 14.VI.1929 , Ψ (OSEC); 14.VI.1949 , ɗ (OSU). Victoria County, Victoria, 1.VIII.1902 , ɗ (USNM); 15.VII.1903 , Ψ (USNM); 12–15.VII.1946 , ɗ (KSUC). Walker County, 8 mi . W of Huntsville, 2.IX.1960 , 2 ɗ (EMEC, UCDC). Wharton County, 7 mi . E Wharton, 9.VIII.1921 , Ψ (EMEC). MEXICO . CHIHUAHUA: Colonia Dublan, July 1931 , ɗ (BYU); Delicias, 13.VII.1947 , 4 Ψ (FSCA); Delicias, 4150', 11.VII.1947 , 3 ɗ, 2 Ψ (FSCA). COAHUILA: 10 mi . S of Ciudad Acuña, 13.IX.1977 , ɗ (EMEC); 10 mi . S of Sabinas, 11.IX.1976 , Ψ (EMEC); Muzquiz, Las Ruscias, 3.VIII.1938 , ɗ (TAMU); Zaragoza, 11.IX.1976 , Ψ (EMEC). NUEVO LEÓN: Monterrey, Chipinque, V.1957 , ɗ (FSCA); 6 mi . S of Monterey, 12.VIII.1959 , Ψ (FSCA). SAN LUIS POTOS: Valles, 1.VII.1959 , Ψ (FSCA).