Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes Author Huang, Jia Author Gong, Lu Author Tsaur, Shun-Chern Author Zhu, Lin Author An, Keying Author Chen, Hongwei text Zootaxa 2019 2019-09-30 4678 1 1 75 journal article 25341 10.11646/zootaxa.4678.1.1 96c8a18a-e887-4345-b266-d76076e8416b 1175-5326 3468782 91B46647-B441-41F5-BE27-3A67D699788C Phortica ( Phortica ) dianzangensis Gong & Chen , sp. nov. ( Fig. 33 ) Diagnosis. Vertical process of gonopod apically small, nearly membraneous, only with 1 projection per side ( Fig. 33D ); 2 pairs of basal bridges of aedeagal median rod fused to each other ( Fig. 33D ). Description. Frons dark brown above, brownish below. Face brownish. Mesoscutum, pleura, and scutellum brownish yellow, with brown to dark brown patches and pollinose pattern. Male terminalia ( Fig. 33 ): Epandrium pubescent anteroventrally, with ca. 10 setae on dorsal to posterolateral portion per side ( Fig. 33B ). Surstylus lacking pubescence on outer surface, with several wedge-like prensisetae on apical margin and inner surface ( Fig. 33C ). Paramere expanded distally; apical process with 2 sensilla and 1 tooth; medial process with 2 sensilla, lacking pubescence or tooth; basal process acute apically ( Fig. 33E ). Aedeagal median rod with 2 slightly asymmetrical processes submedially ( Fig. 33D ). FIGURE 33. Phortica ( Phortica ) dianzangensis Gong & Chen , sp. nov. , male. A. Sixth tergite (lateral view); B. Epandrium and cercus (lateral view); C. Surstylus (frontal view); D. Hypandrium, aedeagus, gonopod, and aedeagal apodeme (lateral view); E. Paramere (lateral view). Scale bars = 0.1 mm. Measurements and indices. BL = 3.78 mm in holotype (range in 7♂ paratypes : 3.33–4.13), ThL = 1.56 mm (1.53–1.87), WL = 2.98 mm (2.67–3.00), WW = 1.24 mm (1.20–1.47), arb = 5/3 (4–5/2–3), avd = 0.58 (0.50–0.80), adf = 2.00 (1.27–2.00), flw = 1.88 (1.45–2.20), FW/HW = 0.42 (0.40–0.56), ch/o = 0.08 (0.08–0.11), prorb = 1.25 (1.00–1.33), rcorb = 0.42 (0.33–0.56), vb = 0.63 (0.41–0.80), dcl = 0.48 (0.36–0.50), presctl = 0.60 (0.50–0.59), sctl = 1.13 (1.00–1.20), sterno = 1.58 (0.80–1.60), orbito = 1.67 (1.13–1.57), dcp = 0.27 (0.24–0.33), sctlp = 1.07 (0.94–1.13), C = 2.12 (1.97–2.96), 4c = 1.50 (1.53–1.78), 4v = 2.64 (2.68–3.27), 5x = 0.93 (0.79–1.00), ac = 3.67 (3.21–3.91), M = 0.59 (0.54–0.71), C3F = 0.68 (0.61–0.74). Type materials. Holotype ♂ ( SCAU , No. 123251), CHINA : Beibeng , Motuo , Xizang , 29°19'00''N , 95°20'00''E , alt. 1000m , 29.ix.2010 , J.J. Gao . Paratypes : CHINA : 1♂ ( TMNS ), same data as the holotype ; 15♂ , Muyiji Park , Ximeng , Yunnan , 22°37'15''N , 99°35'42''E , alt. 1200m , 6♂ ( SCAU , Nos. 123252–55, 112044 , 112045 ) on 4.iv.2011 , J.M. Lu , Z.F. Shao , Y.R. Su , S.J. Yan , 5♂ ( KIZ , Nos. 0090527–31) on 30.iv.2016 , J. Huang , Y.Q. Liu , Y.L. Wang , L. Zhu , 4♂ ( SCAU , Nos. 112046 –49) on 26.x.2017 , H.W. Chen , L. Gong , B.X. LI ; 4♂ ( KIZ , Nos. 0090675–78), Hesong , Menghai , Yunnan , 21°50'17''N , 100°05'48''E , alt. 1700–1900m , 7.v.2012 , H.W. Chen, J.J. Gao; 6♂ , Guan- lei, Mengla , Yunnan , 21°38'39''N , 101°09'52''E , alt. 560–740m , 1♂ ( KIZ , No. 0090532) on 14.xi.2012 , J.J. Gao , 5♂ ( SCAU , Nos. 124892–96) on 21–23.iv.2016 , J. Huang , Y.Q. Liu Y.L. Wang , L. Zhu ; 35♂ ( SCAU , No. 124897, 112054 –87), Jinshan , Menglian , Yunnan , 22°20'06''N , 99°34'34''E , alt. 1040m , 1♂ on 27.iv.2016 , J. Huang, 28♂ on 23–25.x.2017 , H.W. Chen, L. Gong, B.X. Li, 6♂ on 15.iv.2018 , H.W. Chen, L. Gong, Y. Lin; 6♂ ( SCAU , Nos. 124886–91), Likan , Ximeng , Yunnan , 22°39'21''N , 99°36'28''E , alt. 840m , 1.v.2016 , J. Huang , Y.Q. Liu , Y.L. Wang , L. Zhu ; 1♂ ( SCAU , No. 124843), Mengyuan , Mengla , Yunnan , 21°47'29''N , 101°22'59''E , alt. 990m , 7.viii.2016 , L. Gong; 3♂ ( SCAU , Nos. 124898–900), Botanic Garden , Ruili , Yunnan , 24°01'12''N , 97°51'33''E , alt. 1170m , 22.viii.2016 , H.W. Chen, L. Gong, Y.Q. Liu; 1♂ ( SCAU , No. 124712), Nanling ( Shangsongshan ), Lancang , Yun- nan, 22°47'57''N , 99°55'11''E , alt. 1435m , 23.x.2017 , B.X. Li; 2♂ ( SCAU , 124710, 11), Bangdong , Ruili , Yunnan , 24°06'09''N , 97°50'55''E , alt. 1370m , 6.xi.2017 , H.W. Chen, L. Gong; 1♂ ( SCAU , No. 124713), Zhongshan , Mang- shi, Yunnan , 24°07'49''N , 98°31'27''E , alt. 850m , 8.xi.2017 , H.W. Chen. Distribution. China ( Yunnan , Xizang ). Etymology. Toponym, according to the locality of collected specimens from Yunnan and Xizang .