Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 1 7 49 journal article 10.5281/zenodo.5392079 1638-9387 5392079 Glyphiulus semigranulatus n. sp. ( Figs 13 ; 14 ) TYPE MATERIAL. — China . Yunnan Prov. , Meele (= Mile) County, Bai Long Dong Cave (= White Dragon Cave), FIG. 14. — Glyphiulus semigranulatus n. sp. , ♂ paratype: A , antenna; B , gnathochilarium, ventral view; C , legs 1, caudal view; D , leg 3; E , anterior gonopods, caudal view; F , posterior gonopods, caudal view; G , same, frontal view. Scale bar: 0.2 mm. 2.I.1989 , leg. P. Beron , holotype ♂ ( NMNHS ) ; paratypes 1 ♂ , 1 ♀ ( NMNHS ) ; 1 ♀ ( SEM ) ; 1 ♂ ( MNHN GA 037 ). — Yunnan Prov. , Jianshui County , Yan Zi Dong Cave , 10.I.1989 , leg. P. Beron , paratypes 5 ♂♂ , 3 ♀♀ , 4 juv. ( NMNHS ) ; 4 ♂♂ , 2 ♀♀ ( MNHN GA 037 ) ; 4 ♂♂ , 2 ♀♀ ( ZMUM ) ; 2♂♂ , 2 ♀♀ ( ZMUC ) ; 1 ♂ , 1 ♀ ( MCSNV ) ; 1 ♂ ( SEM ) . ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus . DIAGNOSIS. — Differs by a position somewhat intermediate between the most similar congeners G. granulatus and G. paragranulatus n. sp. , i.e. coloration, carinotaxy, antennae, claws, etc., as in G. granulatus , but size, segment shape, leg length and anterior gonopods more like in G. paragranulatus n. sp. DESCRIPTION Length of adults of both sexes 12-24 mm , width 0.8-1.2 mm . Adults of both sexes with 31-64p+5-1a+T; largest juvenile ♂ with 28p+5a+T, largest juvenile ♀ with 27p+5a+T; two smallest juveniles with 15 or 18 haplopodous segments after 6th, plus 2a+T or 1a+T, respectively.Length of holotype about 22 mm , width 1.0 mm, with 45p+3a+T. FIG. 15. — Glyphiulus beroni n. sp. , ♀ paratype: A , anterior part of body, lateral view; B , same, dorsofrontal view; C , middle part of body, dorsal view; D , same, lateral view; E , caudal part of body, dorsal view; F , same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm. Coloration, carinotaxy ( Fig. 13 A-E), antennae ( Figs 13A ; 14A ), gnathochilarium, claws, etc. ( Fig. 14 B-D, F, G) basically as in G. granulatus , but body size (usually much larger), segment shape (slightly compressed laterally) ( Fig. 13C ), telson ( Fig. 13E, F ) with peculiar flaps on paraprocts near hypoproct ( Fig. 13F ), legs (somewhat longer) and anterior gonopods ( Fig. 14E ) more like those of G. paragranulatus n. sp. Adults with 11-15 ocelli. FIG. 16. — Glyphiulus beroni n. sp. , ♂ paratype: A , antenna; B , legs 1, caudal view; C , legs 2, caudal view; D , anterior gonopods, caudal view; E , posterior gonopods, caudal view; F , same, frontal view. Scale bar: 0.2 mm. REMARKS In the absence of any evident troglomorphic traits, this species can only be considered as troglophilic at most. This is supported by G. semigranulatus n. sp. being present in several relatively strongly isolated caves, sometimes in significant numbers.