A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China Author Leu, Marc Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland Author Bucher, Hugo Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland Author Vennemann, Torsten Author Bagherpour, Borhan Author Ji, Cheng Author Brosse, Morgane Author Goudemand, Nicolas text Swiss Journal of Palaeontology 2022 19 2022-11-22 141 1 1 61 http://dx.doi.org/10.1186/s13358-022-00259-x journal article 10.1186/s13358-022-00259-x 1664-2384 PMC9681704 36439694 12000082 Genus NOVISPATHODUS Orchard, 2005 Type species and holotype . Neospathodus abruptus Orchard, 1995 , pp. 118–119 , figs. 3.23–24. (See figure on next page.) Fig. 16 Neogondolellinae , Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P 1 elements if not specifically identified otherwise. A , F , M Discretella ? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella ? n. sp. B;YC16, PIMUZ 39113. C Discretella aff. discreta (MÜller);YC22,PIMUZ 39106. D , S Discretella discreta (MÜller) ; D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard) ;YC41, PIMUZ 39225. G–I , K , L , N , O Discretella pseudodieneri n. sp. ; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127, L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller) ;YC16, PIMUZ 39140. P–R Neospathodus bevelledi n. sp. ; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180 Fig. 16 (See legend on previous page.) Type stratum and locality . Jabral Safra , Oman . Remarks. Te genus Novispathodus was introduced as a new genus with a 15-element apparatus by Orchard (2005) and revised by Goudemand et al. (2012b) based on the swapping of the S 1 and S 2 positions. Additional to the type species ( Nv. abruptus ) and other species whose multi-element apparatus has been reconstructed (e.g. Nv. pingdingshanensis ( Goudemand et al., 2012b ) , Nv. waageni (unpublished)), we tentatively assign also the following species to Novispathodus on the basis that they co-occur with S and M elements that are reminiscent of Novispathodus .