South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini) Author Gordon, Robert D. Northern Plains Entomology PO Box 65 Willow City, ND 58384, USA Author F, Guillermo González Nocedal 6455 La Reina Santiago, Chile Author Hanley, Guy A. Northern Plains Entomology Minot, ND text Insecta Mundi 2020 2020-05-29 2020 766 1 25 journal article 10.5281/zenodo.5353544 1942-1354 5353544 772BDDF2-AD14-4950-B3B3-D6A2C9E403C2 9. Calloeneis signata (Korschefsky) Cryptognatha signata Korschefsky 1936: 299 ; Blackwelder 1945: 449 . Calloeneis signata ; González 2010: 248 . Description. Male . Length 2.0 mm, width 1.6 mm ; dorsal surface shiny except head alutaceous, pronotum with faint microsculpture. Color yellow; head yellow with brown vertex; pronotum with median 1/3 brown; elytron with suture narrowly brown, brown sutural vitta extended posteriorly to apex, then laterally a short distance, basal 1/2 with large, irregular macula, macula with small, narrow, oblique yellow spot laterad of suture, median portion of macula deeply indented with yellow ( Fig. 32 ); venter entirely yellow except basal ventrite with apex of median 1/3 brown. Head with punctures not visible because concealed in alutaceous sculpture; pronotal punctures small, separated by up to twice a diameter; elytral punctures as large as on pronotum, separated by up to twice a diameter; prosternal punctures small, distinct, separated by a diameter or less; mesosternal punctures larger than on prosternum, separated by less than a diameter, metasternal punctures large, separated by up to twice a diameter medially, absent in lateral 1/3; basal abdominal ventrite with coarse punctures in median 1/3 separated by up to twice a diameter, ventrite 2 coarsely punctured in median 2/3, punctures separated by less than a diameter, ventrites 3, 4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head parallel sided, medially depressed, 1.5 times as wide as eye, apex of clypeus weakly curved ( Fig. 33 ); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 2/3 distance to rear margin of ventrite. Genitalia long, penis guide longer than paramere, basal 1/4 wide, apical 3/4 narrow to rounded apex; paramere widened from base to setiferous apex ( Fig. 34, 35 ); penis long, slender, inner arm apically acute, outer arm long, apically rounded ( Fig. 36 ). Female. Similar to male except head dark brown, clypeus yellowish brown; genitalia with spermathecal capsule equal in width throughout, apex of cornu rounded ( Fig. 37 ). Variation. Length 1.7 to 2.1 mm , width 1.5 to 1.6 mm , elytral color pattern highly variable from that described above to having basal macula nearly all dark brown excepting yellow sutural spot. Type locality. Paraguay , Villarica. Type depository. USNM. Geographical distribution. Argentina , Brazil , Guyana , Paraguay , Peru . Specimens examined. 36. Argentina . Buenos Aires ; Calilegua ; Corrientes ; Loreto ; Misiones , Posadas . Brazil . Cantareira , Sao Paulo ; Santana , Sao Paulo . Guyana . Georgetown , Univ. Guyana Campus . Peru . Junin , Chanchamayo , La Merced ; Madre de Dios , Tahuamanu, Inapari ; Pucallpa. ( BMNH , CPGG , MEUT , USNM ) . Remarks. Calloeneis signata is recognized by its distinctive dorsal color pattern in spite of some variability in that pattern. The Korschefsky collection is property of the USNM and a single specimen in that collection is labeled (red paper) “Typus.” However, that specimen cannot be the holotype because it bears the label “Est. Exp. Loreto, 193......Dr. A. Ogloblin” and the published holotype data are “ Paraguay , Villarica.” A single specimen in the Korschefsky collection does bear the latter label but has no type indication.