Predatory midges of the genus Schizohelea Kieffer, 1917 in Europe and North Africa (Diptera: Ceratopogonidae) Author Szadziewski, Ryszard Department of Invertebrate Zoology and Parasitology, Faculty of Biology, University of Gdańsk, Wita Stwosza 59, 80 - 308 Gdańsk, Poland Author Dominiak, Patrycja TromsØ University Museum, UiT - The Arctic University of Norway, NO- 9037 TromsØ, Norway Author Rumišek, Mario Division of Zoology, Faculty of Science, HR- 10000 Zagreb, Croatia Author Neinhuis, Christoph Chair of Botany, Faculty of Biology, Technische Universität Dresden, 01062 Dresden, Germany Author Wanke, Stefan Chair of Botany, Faculty of Biology, Technische Universität Dresden, 01062 Dresden, Germany & Institut für Ökologie, Evolution und Diversität, Goethe-Universität Frankfurt, 60438 Frankfurt am Main, Germany & Abteilung Botanik und molekulare Evolutionsforschung, Senckenberg Gesellschaft für Naturforschung, 60325 Frankfurt am Main, Germany Author Rupp, Thomas Department of Environment and Biodiversity, Paris-Lodron University of Salzburg, Hellbrunnerstr. 34, 5020 Salzburg, Austria text Zootaxa 2024 2024-09-26 5514 2 169 187 http://dx.doi.org/10.11646/zootaxa.5514.2.5 journal article 10.11646/zootaxa.5514.2.5 1175-5326 13849728 B20A1F98-0B55-48DF-AD7F-EA4339591E32 Schizohelea incerta ( Clastrier, 1963 ) ( Figs. 4a–d , 5 ) Monohelea incerta Clastrier, 1963: 60 (female, Algeria ). FIGURE 4. Schizohelea incerta ( Clastrier, 1963 ) (a–d) and S. algeriana Szadziewski, Dominiak, Rumišek et Rupp , sp. nov. (e–h): a—male maxillary palp; b—male wing; c—female wing; d—male thorax and legs; e—male wing; f—female wing; g—female genitalia; h—spermathecae. FIGURE 5. Schizohelea incerta ( Clastrier, 1963 ) , male genitalia: a—ventral view; b—lateral view; c—gonocoxal expansions; d, e—parameres; f–h—aedeagus. Diagnosis . In both sexes second radial cell about 1.2–1.3 times longer than first radial cell. Males are distinct in having aedeagus with two apical submedian processes and a ventral plate armed with strong teeth, slender parameres armed with a short claw-like spine, and bilobed lateroventral tubercle of gonocoxite. Female with or without macrotrichia on posterior half of wings, with unarmed legs, and arms of sternite 9 broad and blunt, each with dark recurved apical margin. Description Male . Body black with slightly paler tarsi ( Fig. 4d ). Flagellum 0.73–0.74 mm long. Third palpal segment 0.070 –0.085 mm long ( Fig. 4a ). Wing length 1.37–1.50 mm , costal ratio CR 0.58–0.64; second radial cell about 1.2–1.3 times longer than first one ( Fig. 4b ). Wing membrane without macrotrichia. Knob of haltere pale with few short setae. Scutellum with 4 bristles and several short setae. Legs slender ( Fig. 4d ), tarsal ratio TR(1) 1.5–1.7, TR(2) 1.8, TR(3) 1.6–1.8. Gonocoxite with blunt lateroventral bilobed tubercle ( Figs. 5a, c ). Gonostylus slender, slightly C-curved, about 1.3 times longer than gonocoxite. Parameres slender with short claw-like dark apical curved spine ( Figs. 5d–e ). Aedeagus arch-shaped with two submedian apical projections and dentate ventral plate ( Figs. 5f–h ). Female . Similar to male. Flagellum length 0.57–0.61 mm , antennal ratio AR 0.98–1.00. Third palpal segment 0.070 mm long. Mandible armed with 10–12 teeth. Wing length 1.23–1.42 mm , costal ratio CR 0.64–0.70; second radial cell about 1.2–1.3 times longer than first radial cell. Wing membrane in two specimens with few macrotrichia at anterior apex ( Fig. 4c ). Scutellum with 4 bristles and several short setae. Tarsal ratio TR(1) 1.7–1.8, TR(2) 1.9–2.2, TR(3) 2.4–2.5. Spermathecae almost spherical, subequal, length with neck 0.053 –0.060 mm . Rudimentary spermatheca indistinct. Sternite 9 separated medially with broad, blunt single arms, each with dark recurved apical margin. Material examined . Tunisia . Jendouba Governorate , suburbia of Tabarka , inside trap-flowers of Aristolochia navicularis , 15.03.2006 , N36 o 55’03.3 ’’ E08 o 45’12’’ , 3 females , 11 males , Ch. Neinhuis & S. Wanke leg. Distribution . Algeria and Tunisia in North Africa. Both sexes visit flowers of Aristolochia navicularis . From Tunisia recorded for the first time. Comments . The male is described and illustrated for the first time.