Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria pacitanensis sp. nov. urn:lsid:zoobank.org:act: E0076893-CF05-4D23-A1E0-402A2DE7F0EF Figs 9 , 41 , 53 , 62 , 65 ; Tables 3–4 Diagnosis Landouria pacitanensis sp. nov. is characterized by a strongly depressed conical, brownish-corneous, sharply keeled shell and a penis with a distinct constriction between its distal and its proximal part, and the presence of a coecum at the proximal end of the penis. Etymology Landouria pacitanensis sp. nov. is named after its occurrence in the Pacitan Regency. Material examined Holotype INDONESIA • holotype (det. anat.); East Java , Pacitan , Srau Beach ; 8°15′01″ S , 110°59′46″ E ; 60 m a.s.l. ; 10 Aug. 2014 ; A.S. Nurinsiyah , F.L.H. Irsyad , F.J. Sari and E. Nurlaela leg.; measurements: D = 12.7 mm , H = 7.1 mm ; MZB 21191 . Figs 63–71 . Genitalia of species of Landouria from Java . 63 . L. tonywhitteni sp. nov. , Central Java , Taruman Village, near Sendang Sedayu ( holotype , MZB 19214). 64 . L. nodifera sp. nov. , Central Java , Larangan Village, Lanang Spring ( holotype , MZB 19227). 65 . L. pacitanensis sp. nov. , East Java , Pacitan, near Srau Beach (MZB 19231). 66 . L. intumescens ( Martens, 1867 ) , East Java , Guwoterus (MZB 19207). 67 . L. smimensis ( Mousson, 1848 ) , East Java , Tosari (ZMA 390881). 68 . L. zonifera sp. nov. , Yogyakarta , Kulonprogo, Sibolong ( holotype , MZB 17727). 69 . L. ciliocincta ( Möllendorff, 1897 ) , East Java , Malang, Kondangmerak (MZB 19211). 70 . L. pakidulan sp. nov. , Central Java , Nusa Kambangan Island, nature monument Gligir ( holotype , ZMA 417011a). 71 . L. menorehensis sp. nov. , Yogyakarta , Kulonprogo, Sibolong ( holotype , MZB 19201). Scale bars: 1 mm . Paratypes INDONESIA – East Java • 7 spec. ; Pacitan , Srau Beach ; 8°14′58″ S , 110°59′42″ E ; 78 m a.s.l. ; MZB 19304 • 4 spec. ; same data as for preceding; ZMH 148511 • 6 spec. ; same data as for holotype; MZB 19234 • 11 spec. ; same data as for holotype; MZB 21270 • 4 spec. ; same data as for holotype; ZMH 148510 • 6 spec. ; same data as for holotype; ZMH 148513 . Other material INDONESIA – East Java • 1 spec. (det.anat.); Pacitan , near Srau Beach ; 8°13′55″ S , 111°00′54″ E ; 217 m a.s.l. ; MZB 21267 • 11 spec. ; same data as for preceding; MZB 19231 • 5 spec. ; same data as for preceding; ZMH 148509 • 4 spec. ; same data as for preceding; ZMH 148512 • 7 spec. ; Pacitan , near Srau Beach ; 8°13′54″ S , 111°00′55″ E ; 223 m a.s.l. ; MZB 21192 • 5 spec. ; same data as for preceding; ZMH 148507 . Description SHELL ( Figs 9 , 62 ; Tables 3–4 ). Depressed conical, with 5.25–6 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture almost circular; upper insertion of peristome descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 28–34% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 41 , 65 ; Table 4 ). Atrium short; penis long, consisting of a long, cylindrical and very muscular distal part covered by an additional sheath, which is connected by muscular tissue to proximal part of epiphallus, and a dilated proximal part; distal and proximal parts of penis separated by a narrow constriction; penial papilla indistinct; inside proximal part of penis with longitudinal pilasters, which pass into a more complex sculpture in distal part of penis; at proximal end of penis with a blunt penis coecum; penial retractor runs from diaphragm to proximal part of epiphallus, which tapers proximally; flagellum moderately long, broadest in its middle part, without node, gradually tapering towards its proximal end; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Remarks Landouria nodifera sp. nov. , L. tonywhitteni sp. nov. , L. pacitanensis sp. nov. , L. sukoliloensis sp. nov. , L. sewuensis sp. nov. and L. intumescens share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. Landouria pacitanensis sp. nov. shares a coecum at the proximal end of the penis with L. nodifera sp. nov. , from which it differs in the long and very muscular distal part of the penis, which is separated from the proximal part of the penis by a very narrow constriction, the insertion of the penial retractor close to the distal end of the epiphallus and the lack of a node at the base of the flagellum. The populations from the surroundings of Srau Beach differ in shell size and shape. The populations closer to the beach are smaller (D 10.6–12.7 mm vs 13.7–14.7 mm ) and more elevated (D/H 1.57–1.83 vs 1.70–2.18) than those from further inland. Distribution Landouria pacitanensis sp. nov. is endemic to the Pacitan Regency in East Java (Fig. 53).