Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae) Author Fehér, Zoltán E801EC76-8B1E-450B-993E-BBBE57C00EA9 Department of Zoology, Hungarian Natural History Museum, Baross u. 13, Budapest, H- 1088, Hungary. feher.zoltan@nhmus.hu Author Deli, Tamás A33DCDF3-6293-4B34-B1DD-330211905D74 Móricz Zsigmond u. 2, Gyomaendrőd, H- 5500, Hungary. virpazaria@gmail.com Author Erőss, Zoltán P. 0B2C74A7-0001-4E25-984C-88CB0C705313 Bem u. 36., Budapest, H- 1151, Hungary. erosspeter@hotmail.com Author Lika, Romilda 9662552B-B203-43FF-B427-4330080E1CBA Departament of Biology, Faculty of Natural Sciences, Tirana University, Bulevardi Zogu i Parë, Tiranë, AL- 1001, Albania. romildalika@gmail.com text European Journal of Taxonomy 2019 2019-09-26 558 1 25 journal article 10.5852/ejt.2019.558 5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59 2118-9773 3474974 5B9DC474-1F26-4732-B7B8-32C285FE0099 Virpazaria (Virpazaria) gittenbergeri Fehér & Erőss sp. nov. urn:lsid:zoobank.org:act: C987B11A-418B-48B2-A5D5-B71966765600 Fig. 4 A–E Spelaeodiscus sp. (juv.) – Subai 2009: 111 . Differential diagnosis This species can be distinguished from congeners by its small size and by its having one knob on the palatal peristome, and from species of the related Spelaeodiscus , which have one knob/tooth on the peristome by its inferiorly broader rounded body whorl and narrower aperture. Etymology This species is dedicated to and named after Edmund Gittenberger (Leiden, the Netherlands ), a prominent malacologist who did pioneering work on the subterranean terrestrial gastropods of the Balkans. The name is used as the genitive of a noun of male gender. Material examined Holotype MONTENEGRORumija Mts , 2 km S of Štegvaš Pass on Ostros–Ulcinj road ( 4 km S of Arbnež ); 42.0582° N , 19.3535° E ; 350 m alt.; 28 May 2015 ; T. Deli , Z. Erőss and Z. Fehér leg.; HNHM 103411 . Paratypes MONTENEGRO20 spec. ; same collection data as for holotype; HNHM 103413 20 spec. ; same collection data as for holotype; NHMW 111671 20 spec. , 38 fr.; same collection data as for holotype; DT 8 ad. , 8 juv. ; same locality as for holotype; 16 Jul. 2017 ; Z. Erőss and Z. Fehér leg.; HNHM 103451 8 ad. , 9 juv. ; same locality as for holotype; 16 Jul. 2017 ; Z. Erőss and Z. Fehér leg.; NHMW 111674 16 ad. , 9 juv. ; same locality as for holotype; 16 Jul. 2017 ; Z. Erőss and Z. Fehér leg.; ER 2 ad. , 1 juv. ; SW of Štegvaš Pass , 2 km S of Arbnež on Ostros–Ulcinj road; 42.0636° N , 19.3710° E ; 480 m alt.; 16 Jul. 2017 ; Z. Erőss and Z. Fehér leg.; HNHM 103445 3 ad. , 1 juv. ; same collection data as for preceding; NHMW 111673 3 ad. , 1 juv. ; same collection data as for preceding; ER . Other material ALBANIAShkodër district , Mt Tarabosh , Vallas , near mosque ; 42.0506° N , 19.4162° E ; 190 m alt.; 27 May 2015 ; T. Deli , Z. Erőss and Z. Fehér leg.; HNHM 103415 same collection data as for preceding; NHMW 110430/MN/0986 same collection data as for preceding; DT . Dimensions Holotype : H 1.55 mm ; W 2.5 mm ; Ha 1.0 mm; Wa 1.1 mm . Paratypes (HNHM 103413, n =5): H 1.4–1.6 mm ; W 2.35–2.5 mm . Vallas population (HNHM 103415, n= 5): H 1.25–1.5 mm ; W 2.2–2.4 mm . Description Shell discoid, spire barely raised, consists of 4¼–4¾ regularly increasing whorls, of which 1½–2 are protoconch whorls. Fresh shells glassy translucent, older shells whitish. Protoconch grained. Teleoconch whorls equidistantly ribbed: 59–69 ribs on body whorl and fine, regular lines in between. Fig. 4. Shells of species of Virpazaria Gittenberger, 1969 . A–E . Virpazaria (Virpazaria) gittenbergeri Fehér & Erőss sp. nov. , holotype (HNHM 103411). F–J . Virpazaria (Virpazaria) pesici Fehér & Deli sp. nov. , holotype (HNHM 103403). Scale bars = 1 mm (note that the latter species is less magnified). In bottom view, umbilicus wide, 1/4–1/5 of total shell width, a small part of it overlaid by columellar part of aperture. In frontal view, silhouette of body whorl asymmetrically rounded (shouldered): broader curved in bottom, more angulated on upper side and near suture horizontally flattened. Suture barely depressed.Aperture narrow crescent-shaped. Instead of a flap, with an outstretched swelling (a thickened and emerged callus) on parietal side. One knob on palatal peristome, ca at one-third of way between its angular and columellar junctions. Peristome somewhat reflected between columellar junction and palatal knob, appearing thickened in frontal view. Above knob, peristome simple, delineating an apertural sinus. In side view, parietal swelling oblique (inferior part stands forward) and aperture sigmoidally inclined (inferior part stands backward). Distribution This species is known from the southeastern part of the Rumija Mts (also known as Mt Tarabosh). The Štegvaš Pass ( Montenegro ) and Vallas ( Albania ) populations are within a distance of 5 km of one another. Both sites are on Upper–Middle Triassic limestone. At the type locality, this species was found together with V. stojaspali and V. pesici . Conservation status To our present knowledge, this is a narrow-range endemic species with an area of occurrence (AOO) smaller than 20 km 2 . However, there is no reason to suppose that AOO, extent of occurrence (EOO), number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT). Remarks The Vallas population differs from the typical form by its denser ribbing (76–88 ribs on the body whorl).