Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)
Author
Fehér, Zoltán
E801EC76-8B1E-450B-993E-BBBE57C00EA9
Department of Zoology, Hungarian Natural History Museum, Baross u. 13, Budapest, H- 1088, Hungary.
feher.zoltan@nhmus.hu
Author
Deli, Tamás
A33DCDF3-6293-4B34-B1DD-330211905D74
Móricz Zsigmond u. 2, Gyomaendrőd, H- 5500, Hungary.
virpazaria@gmail.com
Author
Erőss, Zoltán P.
0B2C74A7-0001-4E25-984C-88CB0C705313
Bem u. 36., Budapest, H- 1151, Hungary.
erosspeter@hotmail.com
Author
Lika, Romilda
9662552B-B203-43FF-B427-4330080E1CBA
Departament of Biology, Faculty of Natural Sciences, Tirana University, Bulevardi Zogu i Parë, Tiranë, AL- 1001, Albania.
romildalika@gmail.com
text
European Journal of Taxonomy
2019
2019-09-26
558
1
25
journal article
10.5852/ejt.2019.558
5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59
2118-9773
3474974
5B9DC474-1F26-4732-B7B8-32C285FE0099
Virpazaria (Virpazaria) gittenbergeri
Fehér & Erőss
sp. nov.
urn:lsid:zoobank.org:act:
C987B11A-418B-48B2-A5D5-B71966765600
Fig. 4
A–E
Spelaeodiscus
sp. (juv.) –
Subai 2009: 111
.
Differential diagnosis
This species can be distinguished from congeners by its small size and by its having one knob on the palatal peristome, and from species of the related
Spelaeodiscus
, which have one knob/tooth on the peristome by its inferiorly broader rounded body whorl and narrower aperture.
Etymology
This species is dedicated to and named after Edmund Gittenberger (Leiden,
the Netherlands
), a prominent malacologist who did pioneering work on the subterranean terrestrial gastropods of the Balkans. The name is used as the genitive of a noun of male gender.
Material examined
Holotype
MONTENEGRO
•
Rumija Mts
,
2 km
S of Štegvaš Pass on Ostros–Ulcinj
road (
4 km
S of Arbnež
);
42.0582° N
,
19.3535° E
;
350 m
alt.;
28 May 2015
;
T. Deli
,
Z. Erőss
and
Z. Fehér
leg.;
HNHM 103411
.
Paratypes
MONTENEGRO
•
20 spec.
; same collection data as for holotype;
HNHM 103413
•
20 spec.
; same collection data as for holotype;
NHMW
111671
•
20 spec.
, 38 fr.; same collection data as for holotype;
DT
•
8 ad.
,
8 juv.
; same locality as for holotype;
16 Jul. 2017
;
Z. Erőss
and
Z. Fehér
leg.;
HNHM 103451
•
8 ad.
,
9 juv.
; same locality as for holotype;
16 Jul. 2017
;
Z. Erőss
and
Z. Fehér
leg.;
NHMW
111674
•
16 ad.
,
9 juv.
; same locality as for holotype;
16 Jul. 2017
;
Z. Erőss
and
Z. Fehér
leg.;
ER
•
2 ad.
,
1 juv.
;
SW of Štegvaš Pass
,
2 km
S of Arbnež on Ostros–Ulcinj
road;
42.0636° N
,
19.3710° E
;
480 m
alt.;
16 Jul. 2017
;
Z. Erőss
and
Z. Fehér
leg.;
HNHM 103445
•
3 ad.
,
1 juv.
; same collection data as for preceding;
NHMW
111673
•
3 ad.
,
1 juv.
; same collection data as for preceding;
ER
.
Other material
ALBANIA
•
Shkodër district
,
Mt Tarabosh
,
Vallas
,
near mosque
;
42.0506° N
,
19.4162° E
;
190 m
alt.;
27 May 2015
;
T. Deli
,
Z. Erőss
and
Z. Fehér
leg.;
HNHM 103415
•
same collection data as for preceding;
NHMW
110430/MN/0986
•
same collection data as for preceding;
DT
.
Dimensions
Holotype
: H
1.55 mm
; W
2.5 mm
; Ha 1.0 mm; Wa
1.1 mm
.
Paratypes
(HNHM 103413, n =5): H
1.4–1.6 mm
; W
2.35–2.5 mm
. Vallas population (HNHM 103415, n= 5): H
1.25–1.5 mm
; W
2.2–2.4 mm
.
Description
Shell discoid, spire barely raised, consists of 4¼–4¾ regularly increasing whorls, of which 1½–2 are protoconch whorls. Fresh shells glassy translucent, older shells whitish. Protoconch grained. Teleoconch whorls equidistantly ribbed: 59–69 ribs on body whorl and fine, regular lines in between.
Fig. 4.
Shells of species of
Virpazaria
Gittenberger, 1969
.
A–E
.
Virpazaria (Virpazaria) gittenbergeri
Fehér & Erőss
sp. nov.
, holotype (HNHM 103411).
F–J
.
Virpazaria (Virpazaria) pesici
Fehér & Deli
sp. nov.
, holotype (HNHM 103403). Scale bars = 1 mm (note that the latter species is less magnified).
In bottom view, umbilicus wide, 1/4–1/5 of total shell width, a small part of it overlaid by columellar part of aperture. In frontal view, silhouette of body whorl asymmetrically rounded (shouldered): broader curved in bottom, more angulated on upper side and near suture horizontally flattened. Suture barely depressed.Aperture narrow crescent-shaped. Instead of a flap, with an outstretched swelling (a thickened and emerged callus) on parietal side. One knob on palatal peristome, ca at one-third of way between its angular and columellar junctions. Peristome somewhat reflected between columellar junction and palatal knob, appearing thickened in frontal view. Above knob, peristome simple, delineating an apertural sinus. In side view, parietal swelling oblique (inferior part stands forward) and aperture sigmoidally inclined (inferior part stands backward).
Distribution
This species is known from the southeastern part of the Rumija Mts (also known as Mt Tarabosh). The Štegvaš Pass (
Montenegro
) and Vallas (
Albania
) populations are within a distance of
5 km
of one another. Both sites are on Upper–Middle Triassic limestone. At the
type
locality, this species was found together with
V. stojaspali
and
V. pesici
.
Conservation status
To our present knowledge, this is a narrow-range endemic species with an area of occurrence (AOO) smaller than
20 km
2
. However, there is no reason to suppose that AOO, extent of occurrence (EOO), number of locations, number of subpopulations or the number of mature individuals are declining or extremely fluctuating. Therefore, it might be assessed as Near Threatened (NT).
Remarks
The Vallas population differs from the typical form by its denser ribbing (76–88 ribs on the body whorl).