Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae) Author Martens, Jochen Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany; Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany Author Maghradze, Eter Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia Author Barjadze, Shalva Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia text Zootaxa 2021 2021-04-07 4951 3 541 558 journal article 7383 10.11646/zootaxa.4951.3.7 a6ee038f-4cda-4f38-8fde-8ad0efec8f0f 1175-5326 4668453 EB876346-4282-4B4E-8585-ED50F1A616E4 Nemastomatidae in the Holarctic This opilionid family of predominantly small species shows a holarctic distribution. The nominate subfamily, Nemastomatinae Simon, 1879 has its strongholds in the West Palearctic with a distributional focus in central and southern Europe, Turkey and the Caucasus ( Schönhofer 2013 ); a few recently discovered outliers extend the area to Central Asia and western China ( Snegovaya 2010 ; Martens 2016 , 2017 ; Zhang & Martens 2018 ). The subfamily comprises about 120 species ( Schönhofer 2013 ; Martens 2016 ). The second subfamily, Ortholasmatinae Shear & Gruber, 1983 ( Shear & Gruber 1983 ; Shear 2006 , 2010 ) is distributed in East Asia including Japan ( Martens 2019 ; Schwendinger & Gruber 1992 ), and the west coast of North America. Its area extends southward to Central America, the Mexican highlands and the Honduran tropics ( Shear 2010 ; Cruz-Lopéz et al. 2018 ). This subfamily is much poorer in genera and species than Nemastomatinae ; seven genera of the former are described and 18 of the latter. The majority of Ortholasmatinae species occur in western North America and in Central America ( Martens 2019 ).