<p> <strong> A revision of Afrotropical Quasimodo flies (Diptera: Schizophora; Curtonotidae). Part IV — the continental Afrotropical species of <em> Curtonotum </ em> Macquart, with descriptions of thirteen new species and a combined phylogenetic analysis of the Curtonotidae </ strong> </ p> Author Kirk-Spriggs, Ashley H. ashley.kirk-spriggs@nasmus.co.za Author Wiegmann, Brian M. ashley.kirk-spriggs@nasmus.co.za text Zootaxa 2013 2013-07-09 3684 1 1 166 http://dx.doi.org/10.11646/zootaxa.3684.1.1 journal article 10.11646/zootaxa.3684.1.1 1175-5326 5298914 E922034E-1247-400B-97F6-1778CF766B91 Curtonotum pauliani Tsacas, 1974 Figs 127 , 151 , 188 , 286, 289, 292 , 331 . Curtonotum pauliani : Tsacas, 1974: 715 ; figs 7a–d (p. 716), fig. 8b (p. 718). Type locality: [ Madagascar ] “Nosy Mitsio”. Curtonotum pauliani : Kirk-Spriggs (2008c: 246 , fig. 6, 251). Curtonotum pauliani : Kirk-Spriggs (2011) ; figs 12, 25, 38, 69, 72, 75, 90, 103. Remarks : This species was redescribed and figured by Kirk-Spriggs (2011) , only figures are included here for comparative purposes: head and thorax ( Fig. 127 ); frons ( Fig. 151 ); wing ( Fig. 188 ); male terminalia ( Figs 286, 289, 292 ). Differential diagnosis . Curtonotum pauliani belongs to a group of ten species here ascribed to the uncinatum species-group that are virtually identical externally and are only separable based on minor differences in the male phallus (see Differential diagnosis under C . uncinatum sp. n. for details). Curtonotum pauliani can be separated from other species in the species-group by reference to the above key. Type material examined : MADAGASCAR : holotype ♂ , “ Madagascar Nord-Ouest / Nosy Mitsio / 13– 14.I.[19]60 / R. Paulian // INSTITUT / SCIENTIFIQUE / MADAGASCAR [pale grey card] // HOLOTYPE [red card] // CURTONOTUM / pauliani / Holotype ♂ n.sp. / L. TSACAS DET. 1973 [printed & handwritten] // MUSÉUM PARIS // Curtonotum / pauliani ♂ / Tsacas, 1974 / A.H. Kirk-Spriggs vidit 2006” [head missing] (MNHN). In fair condition, head, right fore leg, left fore tarsus and hind leg missing; re-staged on nu-poly mount; dissected, abdomen and terminalia in micro-vial pinned beneath specimen. Paratypes (all labelled: “ PARATYPE [red card] // CURTONOTUM / pauliani / n.sp. / L. TSACAS DET. 1973 [printed & handwritten] // Curtonotum / pauliani ♂ [or ♀ ] / Tsacas, 1974 / A.H. Kirk-Spriggs vidit 2006”): MADAGASCAR : 1♀ , same labels as holotype , except: “ ALLOTYPE [red card] // CURTONOTUM / pauliani / n.sp. / L. TSACAS DET. 1973 [printed & handwritten]” [wing detached, glued to card]; 2♂ , same labels as holotype [1 head missing]; 1♀ , same labels as holotype , except: “EXEMP. / DESS. [printed; orange card]”; 1♀ , same labels as holotype , except: “ouile gameha / ma tres large / Anomalié ♀ [handwritten]”; 1♂ , “ Madagascar Nord-Ouest / dct. Majunga / forêt Ankarafantsitka [= Ankarafantsika] 120m / XII-[19]59 / Raharizonina // INSTITUT / SCIENTIFIQUE / MADAGASCAR [pale grey card] // MUSÉUM PARIS” [head missing]; 1♂ , “MUSEUM PARIS / MADAGASCAR / PROV. D’ANALALAVA / MAROMANDIA / R. DECARY 1922 [grey card] // PARATYPE [red card] // Specimen / missing from mount / 2006”; 1♂ , 1♀ , same labels, except: “Pr A18 / ♂ [handwritten] [head missing]” (all MNHN). Additional material examined (both labelled “ Curtonotum pauliani Tsacas, 1974 , det. A.H. Kirk-Spriggs 2011 ”): NAMIBIA : 1♂ , Namibia : OPUWA DIST., Ekuju village : Kunene R., 17°19'30"S 13°48'56"E , 11– 12.x.1999 , [ A .H.] Kirk-Spriggs , [T.] Pape , [W.] Hauwanga , Malaise traps , riverine forest ( NMNW ) . SOUTH AFRICA : 1♂ , 6664, SO. AFRICA: Natal , 15 km SE Rorke’s Drift , 28°30'S , 30°30'E , 30.v.1982 , R.M. Miller ( NMSA ) . Distribution . Madagascar , Namibia and South Africa ( Fig. 331 ). The only species known to occur in the continental Afrotropical Region and on the island of Madagascar . Bionomics . In southern Africa the species occurs in the Nama Karoo and Drakensburg Afromontane Grassland and Woodland major habitat types ; Deserts and Xeric Shrublands and Montane Grasslands vegetation types (Appendix III ). This species has been sampled in a Malaise trap in riverine forest in Namibia . A detailed account of habitat associations in Madagascar was provided by Kirk-Spriggs (2011) .