A review of the genus Stilbocistela (Coleoptera: Tenebrionidae: Alleculinae) with description of new species from Malaysia and Indonesia Author Novák, Vladimír text Acta Entomologica Musei Nationalis Pragae 2009 2009-12-15 49 2 783 794 journal article 10.5281/zenodo.5324860 0374-1036 5324860 Stilbocistela jelineki sp. nov. ( Figs. 1–5 ) Type locality. Indonesia , Irian Jaya , Puncak Jaya Pass. Type material. HOLOTYPE : J, ‘ INDONESIA Irian Jaya / Puncak Jaya Pass / 22.i.2002 ; J. Doe lgt.’ ( NMPC ) . PARATYPE : 1 ♀ , same data as holotype ( VNPC ). The types are provided with a printed red label: ‘ Stilbocistela jelineki sp. nov. HOLOTYPUS [or PARATYPUS ] V. Novák det. 2008’. Description. Habitus of male holotype as in Fig. 1 , body oval, strongly vaulted. General colour from pale brown to dark blackish brown; elytra with three large reddish-brown spots. Body length 5.59 mm , width at elytral midlength 2.58 mm . Male ( holotype ). Head ( Fig. 2 ) large, posterior part dark brown, shiny, without setation, anterior part before eyes with sparse and short light setation; clypeus pale brown with longer light setation. Punctuation of posterior part indistinct, surface shiny, punctuation of anterior part (before eyes and clypeus) distinct with large, dense and coarse punctures, interspaces between punctures narrow and shiny, interspaces of clypeus with distinct microgranulation. Eyes dark in colour, large, transverse, emarginate, with distinct narrow interspace between eyes. Distance between eyes distinctly longer than length of antennomere 2 and shorter than length of antennomere 3. Ocular index equal to 12.59. Head widest across eyes 1.08 mm , approximately 0.54 times as wide as pronotal base. Visible part of head 0.99 mm long. Antenna ( Fig. 3 ). Relatively short, 2.88 mm long, reaching 0.52 of body length, antennomeres brown, antennomeres 1 and 2, anterior half of antennomere 3 and apical part of antennomere 11 pale brown, antennomeres 4–11 with short and dense brown setation and distinctly rugose, with relatively shallow, dense and light punctures. Antennomeres 4–10 distinctly serrate, antennomere 2 shortest, antennomere 3 distinctly shorter than antennomeres 4–10. Antennomeres 1–3 slightly shiny. Ratios of relative lengths of antennomeres 1–11 equal to 0.69: 0.46: 1.00: 1.13: 1.27: 1.24: 1.33: 1.48: 1.35: 1.49: 1.38. Length / maximum width ratios of antennomeres 1–11 equal to 1.76: 1.15: 2.21: 1.64: 1.84: 1.73: 1.93: 1.91: 1.79: 2.08: 2.13. Maxillary palpus pale brown, concolorous with clypeus and antennomeres 1–3, with light setation and microgranulation, dull; second maxillary palpomere longer than penultimate palpomere, distinctly broadest at apex, penultimate palpomere shortest, ultimate palpomere triangular, both distinctly broader at apex. Ratios of relative lengths of palpomeres 2–4 equal to 1.91: 1.00: 2.70. Length / maximum width ratios of palpomeres 2–4 equal to 2.67: 1.24: 1.03. Pronotum ( Fig. 2 ) reddish-brown, strongly shiny, glabrous, broad and strongly transverse, longest in middle (length 0.90 mm ) and widest at base (width 2.01 mm ); pronotal index equal to 44.99. Border almost complete, only in middle of anterior margin and in one third of posterior margin near posterior angles indistinct; posterior margin bisinuate. Posterior angles slightly obtuse, lateral margins straight in posterior half and slightly rounded anteriorly. Anterior angles conspicuous, distinctly obtuse. Surface sparsely and shallowly punctate, punctures small, interspaces very broad, with microgranulation, strongly shiny. Elytra oval, glabrous, strongly shiny, bicoloured, dark blackish brown with three large reddish-brown spots. Anterior part of each elytron with two spots between interval 2 and 7 and a third spot extending from elytral suture to elytral interval 5 on each side. Elytra 3.70 mm long and 2.58 mm wide, slightly broader than pronotum, widest approximately at midlength. Elytral length / maximum width ratio equal to 1.44. Surface punctate, strial punctures separated by less than one diameter. Elytral intervals with sparse, small punctures and microgranulation, strongly shiny. Epipleura well developed, reddish brown, evenly narrowing in posterior half, in apical half before abdominal sternite 5 parallel-sided and then narrowing to rounded apex. Scutellum large, glabrous, reddish brown, concolorous with pronotum. Legs unicolorous, pale brown, tarsi with dense and long light setation. Femora thicker than tibiae. Tibiae very narrow, slightly dilated at apex with rows of stronger and shorter dark setae. Tarsomeres of all tarsi narrow. Anterior and middle tarsomeres 3 and 4 and posterior tarsomeres 2 and 3 lobed. Ratios of relative lengths of tarsomeres 1–5 and 1–4 equal to 1.00: 0.77: 0.66: 0.98: 1.98 (protarsus), 1.00: 0.39: 0.60: 0.50: 1.04 (mesotarsus), and 1.00: 0.35: 0.27: 0.50 (metatarsus). Both anterior tarsal claws with four teeth. Ventral side of body brown. Abdominal ventrites glabrous, shiny, with microgranulation and shallow logitudinal striae. Aedeagus ( Figs. 4–5 ) pale yellowish brown, distinctly shiny. Basal piece 5.62 times as long as apical piece, basal piece narrow laterally and almost straight, very slightly regularly narrowing dorsally from base to apex of basal piece. Apical piece almost straight and apex rounded anteriorly from dorsal view. Female. Habitus similar to male. Length 5.98 mm ; length of head 0.92 mm ; width of head 1.05 mm . Ocular index equal to 16.33. Pronotal length (along midline) 1.00 mm; pronotal width at base 2.12 mm . Pronotal index equal to 47.25. Elytral length 4.06 mm ; elytral width 2.68 mm . Ratios of relative lengths of antennomeres 1–11 equal to 0.63: 0.57: 1.00: 1.20: 1.13: 1.17: 1.24: 1.33: 1.24: 1.17: 1.48. Length / maximum width ratios of antennomeres 1–11 equal to 1.54: 1.32: 2.69: 2.10: 1.83: 1.77: 1.84: 1.87: 1.75: 1.60: 2.35. Ratios of relative lengths of tarsomeres 1–5 and 1–4 equal to 1.00: 0.51: 0.58: 1.01: 2.04 (protarsus), 1.00: 0.34: 0.31: 0.48: 1.04 (mesotarsus), and 1.00: 0.27: 0.29: 0.54 (metatarsus). Differential diagnosis. Stilbocistela jelineki sp. nov. differs from other species of the genus mainly by the bicoloured elytra (dark brown with reddish-brown spots) in contrast to the more or less unicoloured elytra without reddish-brown spots in the remaining species. In addition, S . jelineki sp. nov. differs from S. malaica sp. nov. by having antennomeres 4–10 each longer than antennomere 3, from S . manoiensis by antennomere 3 almost twice as long as antennomere 2 and from S . luzonica and S . rostislavi sp. nov. mainly by the distance between eyes distinctly shorter than antennomere 3. For further details see the key above. Etymology. This species is dedicated to Josef Jelínek, a renowned specialist on the beetle family Nitidulidae . Distribution. Indonesia : Irian Jaya .