New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber
Author
Brazidec, Manuel
4EAB750B-E20A-4815-8A9F-7FFB10281485
Univ. Rennes, CNRS, Géosciences Rennes, UMR 6118, 35000, Rennes, France.
manuel.brazidec@gmail.com
Author
Vilhelmsen, Lars
96FC3783-9FA7-421E-B292-6718A3762D45
Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Zoological Museum, Universitetsparken 15, DK- 2100, Denmark.
lbvilhelmsen@snm.ku.dk
text
European Journal of Taxonomy
2022
2022-04-13
813
57
86
http://dx.doi.org/10.5852/ejt.2022.813.1733
journal article
55899
10.5852/ejt.2022.813.1733
f7030f8d-22f5-45e1-8291-bbd966e086f9
2118-9773
6468167
5246FAA3-4E32-4923-A4FD-27FAF6B8EF34
Doliopria baltica
sp. nov.
urn:lsid:zoobank.org:act:
36E9B60D-6971-454F-AD9F-DD69D430C16B
Figs 1G
,
6B–D
,
Table 1
Diagnosis
Head globular, with sparse long hairs on vertex; apical segment of maxillary palpi bearing one long seta (
Fig. 6C
); antenna 11-segmented; scape as long as pedicel + F1–F5 combined, scape with apical rim slightly notched; pedicel cylindrical, elongate; F1 narrower; F2–F6 widening; F7–F9 forming distinct club; F7–F8 two times as wide as long; F9 as wide as long (
Fig. 6B
); anterior scutellar pit suboval, not divided; scutellum subquadrate; propodeum punctuate and carinate (
Fig. 6D
); hind wing bordered with long setae along posterior margin and distal part of fore margin; petiole cylindrical, slightly longer than wide; gaster ellipsoidal; tergite 2 with anterior margin excised (
Fig. 6D
).
Etymology
The species name refers to the origin of the amber piece containing the specimen. The specific epithet is to be treated as an adjective.
Type material
Holotype
NHMD-608374
, a complete
female
partially hidden by a milky coat.
Locality and horizon
Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma.
Description
Female
BODY. BL =
1.20 mm
. Body smooth and shiny, sparsely pubescent, hairs very long. Head globular, slightly higher than long with hairs posteriorly (HeL =
0.23 mm
); eye oval, glabrous; maxillary palpi 5-segmented with one long seta on apical segment; antenna 11-segmented inserted on transverse shelf of low elevation; toruli separated by shallow cleft; scape as long as pedicel + F1–F5 combined, as long as head, with apical circumference slightly notched; pedicel as wide as scape, cylindrical; F1 narrower than pedicel and longer than wide; F2–F5 slightly shortening and widening; F6 two times as wide as long; F7–F9 wider than long, longer than previous flagellomeres, forming clava; F9 ovoid (antennomeres length of
holotype
, in mm: Sc-0.19; P-0.05; F1-0.03; F2-0.02; F3-0.02; F4-0.03; F5-0.03; F6-0.03; F7- 0.07; F8-0.07; F9-0.10); mandibles short, only crossing at tips.
Fig. 6. A
.
Basalys villumi
sp. nov.
, paratype (NHMD-608369), habitus in lateral view.
B–D
.
Doliopria baltica
sp. nov.
, holotype (NHMD-608374).
B
. Habitus in lateral view.
C
. Head in lateral view.
D
. Mesosoma and metasoma in lateral view.
E–H
.
Spilomicrus succinalis
sp. nov.
E–F
. Holotype (NHMD-607131).
E
. Habitus in ventrolateral view.
F
. Head and mesosoma in dorsal view.
G
. Paratype (NHMD-608344), habitus in lateral view.
H
. Paratype (NHMD-608354), habitus in lateral view. Abbreviations: asp = anterior scutellar pit; te = tergite excision. Scale bars: A, E–H = 1 mm; B = 0.5 mm; C–D = 0.25 mm.
MESOSOMA. Slightly shorter than metasoma (MsL =
0.43 mm
); pronotum with lateral posterior margin straight; mesoscutum large and slightly convex, without notauli or any other sulci; anterior scutellar pit deep, not divided, suboval; scutellum subquadrate; propodeum minutely punctuate in anterior part and carinate. Fore wing extending beyond metasoma (FwL =
0.77 mm
), micropubescent and bordered with short setae; only Sc+R present, slightly separate from anterior margin of wing, distally ending in slight thickening of marginal vein. Hind wing length two thirds of length of fore wing (HwL =
0.55 mm
), micropubescent; with long setae along posterior margin and distal part of costal margin; three hamuli present; basal cell open. Legs slender, with only hind coxa and femur stouter; tibial spur formula 1-2-2; tarsal claws simple.
METASOMA. Petiole cylindrical, narrow, longer than wide (PtL =
0.10 mm
; PtW =
0.06 mm
), longitudinally striated, bearing several long hairs; gaster ellipsoidal (GL =
0.44 mm
; GH =
0.21 mm
), not sharply pointed at apex, smooth and glabrous; T2 and S2 longest; T2 covering at least two thirds of gaster, its anterior margin medially excised for distance subequal to petiole length; ovipositor slightly exerted.
Male
Unknown.
Comments
In
Masner & García’s (2002)
keys to the genera of
Diapriinae
,
Doliopria baltica
sp. nov.
keys out to
Doliopria
Kieffer, 1910
because of the following characters: frons unarmed, antenna 11-segmented, notauli absent, wings fully developed, Sc+R separate from anterior margin of wing, basal vein absent. The description of the specimen is consistent with the original diagnosis of the genus by
Kieffer (1910)
and the revised diagnosis of
Masner & García (2002)
.
Doliopria baltica
differs from extant representatives of this genus as follow:
D. americana
Fouts, 1926
has a two-segmented club (
Fouts 1926
);
D. brachyptera
Ogloblin, 1960
has F9 more lengthened and is brachypterous;
D. foersteri
Ogloblin, 1960
has short hairs on eyes, flagellomeres longer than wide and lacks the anterior scutellar pit;
D. equatoriana
Ogloblin, 1960
has F2–F5 longer than wide (Oglobin 1960);
D. collegii
Ferrière, 1929
has F9 longer (
Loiácono
et al
. 2013
);
D. flavipes
Kieffer, 1910
has F5–6 as long as wide and the petiole shorter (
Kieffer 1910
);
D. myrmecobia
Kieffer, 1921
has a two-segmented club and the apical flagellomere sharply pointed at the apex (
Loiácono
et al
. 2013
).