Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)
Author
Kluge, Nikita J.
text
Zootaxa
2023
2023-09-08
5343
3
243
272
http://dx.doi.org/10.11646/zootaxa.5343.3.2
journal article
10.11646/zootaxa.5343.3.2
1175-5326
8334181
BB852CA9-6C6E-4016-9972-AAF2C378E41F
Status of
Securiops
.
Kaltenbach
et al
. (2023)
treated
Securiops
as a genus but not as a subgenus, reporting the following its characters as generic ones: ́(1) labium with strongly reduced glossae, enlarged paraglossae, and very broad, hatchet-like palps; (2) tergalii I–IV with two lamellae; (3) legs elongate, with relatively few short setae on dorsal and ventral margins; (4) claws very elongate, without denticles; and (5) lateral margins of posterior abdominal segments with sharp spines». Actually, only the first of these characters constitutes autapomorphy of the subgenus
Securiops
, while the characters (2)–(5) are diagnostic for the taxon
Cloeon
/fg1, to which the subgenus
Securiops
belongs. The regular lateral row of greatly enlarged spines on distal part of cercus (
Figs 71–72
,
105
,
124–125
) is the most important diagnostic character of
Procloeon
s. l.
, to which the subgenus
Securiops
belongs.
Distribution.
Paleotropical (Afrotropical and Oriental Regions).
Species composition.
Afrotropical species:
Procloeon
(
Securiops
)
falcatum
(
Crass 1947
)
comb. n.
;
P.
(
S.
)
macafertiorum
(Lugo-Ortiz in
Lugo-Ortiz & McCafferty 1996
);
P.
(
S.
)
mandrare
(
Jacobus, McCafferty & Gattolliat 2006
)
comb. n.
;
P.
(
S.
)
megapalpus
(
Jacobus, McCafferty & Gattolliat 2006
)
comb. n.
;
P.
(
S.
)
mutadens
(
Jacobus, McCafferty & Gattolliat 2006
)
comb. n.
. Oriental species:
Procloeon
(
Securiops
)
primasia
(
Kaltenbach
et al
. 2023
)
comb. n.
Key to larvae
(the number in parentheses refers to the antithesis)
1(4) Hind protoptera present (
Figs 29
,
67
) (can be poorly visible, since cuticle of hind protopteron is colorless)............... 2
2(3) Abdominal terga IV–X with numerous scales (
Figs 55–56
); terga IV and VII without medial longitudinal stripe (
Fig. 26
)...........................................................................................
P.
(
S.
)
falcatum
3(2) All abdominal terga without scales, with numerous denticles (
Fig. 74
); terga IV–V and VII–IX with contrasting medial longitudinal stripe on light background (
Figs 66, 69
)........................................
P.
(
S.
)
macafertiorum
4(1) Hind protoptera absent (
Fig. 108, 111
) or represented by short vestiges (
Fig. 102
).
5(6) Spines on lateral side of cercus sharply differentiated: in proximal portion of cercus all spines stout, conic and not longer than next cercomere; in distal portion of cercus all spines delicate, widened from base, as long as 2–3 next cercomeres (
Figs 124–125
). Tergalii with apex stretched and pointed, resembling linden leaf (
Figs 18–24
).................
P.
(
S.
)
primasia
6(5) Spines on lateral side of cercus gradually change from shorter in proximal part of cercus to longer in its distal part (
Fig. 105
). Tergalii with apex rounded (
Gillies 1988
: figs 14–19).....................................................................................................
P.
(
S.
)
mutadens
; possibly, also
(
S.
)
mandrare
and
P.
(
S.
)
megapalpus
.