Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae) Author Génier, François Author Moretto, Philippe text Zootaxa 2017 4248 1 1 110 journal article 33324 10.5281/zenodo.439444 cf486ccc-d298-4179-8ccf-e29a9c894ab4 1175-5326 439444 71F0AC03-C1FB-40AB-8532-99A638FC91E9 Digitonthophagus falciger species group Diagnosis . A single species with FLP sclerite right preapical edge deeply notched, basal edge of emargination produced into a sickle-shape projection ( Fig. 150 ); axial sclerite apical portion thong shape, longer than subaxial sclerite.