New species of Pseudoscorpiones (Arachnida) from tree hollows in a Mediterranean oak forest in Spain
Author
Hernández-Corral, Jesús
Author
Zaragoza, Juan A.
Author
Micó, Estefanía
text
Zootaxa
2018
2018-10-09
4497
2
201
225
journal article
29214
10.11646/zootaxa.4497.2.3
138be9ad-b476-48af-93ee-6fd78c04484d
1175-5326
1452117
0C10BF04-D9FD-49D0-BDB5-647DD690A30F
Neobisium (Neobisium) maroccanum
Beier, 1930
(
Figs 7‒10
)
Type
material examined.
Female
paratype
(
NHMW
,
Reg. No.
26648),
Morocco
, Korifla,
24.XI.1924
.
Non-type material examined.
2 ♀
, 2 tritonymphs (
NHMW
,
Reg. No.
26649),
Morocco
,
Tangier-Tetouan
region,
Chaouen province
,
Xauen
,
30.III.1959
, leg.
H. Franz. Identified
by
M. Beier.
Short description (
♀
).
Female
paratype
in extremely bad condition; the few data obtained for this specimen are given in square brackets after those of the females from Xauen, when different. Carapace subquadrate,
chaetotaxy: 22‒23 setae, formula: 4:6:6:6‒7 [6 posterior margin], epistome prominent, equilateral or isosceles triangle-shaped, length 0.025‒0.033, width
0.028‒0.033 mm
[length 0.025, width
0.020 mm
]. Chelicera with 7‒8 setae on hand, fixed finger with 12‒13 teeth, movable finger with 10 [11] teeth, the 2 [3] distal smaller than others, which are of approximately the same size; spinneret long and rounded hyaline tubercle with about 7 [5] silk ducts (
Figs 7, 8
). Tergites I‒V: 6:8:12:11:12 setae. Anterior process of coxa I with simple tooth shape, apically pointed, medial process sclerotized, distinctly prominent, with numerous denticles (
Fig. 10
) [almost straight, with few denticles (
Fig. 9
)]. Pedipalp smooth, trochanter with one strong tubercle on antiaxial face, femur with low tubercle slightly proximad of middle of antiaxial face, four glandular pores on antiaxial face of hand, close to fingers. Distal opening of patella (X) longer than paraxial margin (Y). Fixed chelal finger with 70 mostly pointed and distinctly long teeth, the 10 basal teeth smaller and rounded; dental row of movable chelal finger with 60 teeth, reaching to halfway between trichobothria
b
and
sb
, progressively decreasing in size and flattening after the 20th distal tooth, basal teeth very flattened but still recognizable; two separate sensilla (
p
1 and
p
2) between trichobothria
sb
and
st
, both slightly closer to
st
; lyrifissure
ma1
distad of trichobothrium
sb
, and lyrifissure
ma2
level with
b
. Trichobothria: distance between
ib
and
ist
1.8 [1.7] times that between
ist
and
it
and equal to that between
is
t and finger tip;
est-etit
forming a group in distal third of finger;
it
slightly distad of
et
and closer to finger tip than to
ist
; distance between
st
and
sb
1.4 times that between
sb
and
b
.
FIGURES 7–13. FIGURES 7–10.
Neobisium (N.) maroccanum
Beier, 1930
. 7, distal end of movable finger of left chelicera of paratype female from Korifla; 9, anterior and medial processes of right coxa I,
idem
; 8, distal end of movable finger of right chelicera, female from Xauen; 10, anterior and medial processes of left coxa I,
idem
.
FIGURES 11–13.
Neobisium (N.) hispanicum
n. sp.
, male holotype. 11, left chelicera; 12, distal end of movable finger of left chelicera; 13, anterior and medial processes of left coxa I.
Measurements and ratios
. Female
paratype
from Korifla: carapace 0.66/0.66 (1.0); pedipalp: trochanter 0.44/ 0.22 (2.0), femur 0.92/0.22 (4.2), patella 0.63/0.26 (2.4), hand+ 0.73/0.39 (1.9), hand - 0.63 (1.6), movable finger 0.81, chela+ 1.46/0.39 (3.7), chela - 1.36 (3.5); ratio movable finger/hand+ 1.1, femur/movable finger 1.1, femur/ carapace 1.4, femur/patella 1.5, hand+/patella 1.2, patella X/Y 1.1, chela+/carapace 2.2, chela+/femur 1.6. Females from Xauen: Carapace 0.68‒080/0.65‒0.77 (1.0). Pedipalp: trochanter 0.47‒0.56/0.23‒0.28 (2.0), femur 0.92‒ 1.10/0.23‒0.26 (4.1‒4.2), patella 0.61‒0.74/0.26‒0.33 (2.2‒2.3), patellar club 0.40‒49 (1.5), hand+ 0.70‒0.88/ 0.43‒0.56 (1.6), hand - 0.61‒0.79 (1.4), movable finger 0.85‒0.99, chela+ 1.52‒1.80/0.56 (3.2‒3.5), chela - 1.43‒1.72 (3.1‒3.3); ratio movable finger/hand+ 1.1‒1.2, femur/movable finger 1.1, femur/carapace 1.4, femur/patella 1.5, hand+/patella 1.1‒1.2, patella X/Y 1.1, chela+/carapace 2.2‒2.3, chela+/femur 1.6‒1.7.
Remarks.
Callaini (1988)
pointed out differences between material from Ketama,
Morocco
that he had studied and Beier’s redescription (1963) of
Neobisium maroccanum
. The specimens from Xauen studied here coincide with Callaini’s description (1988) and are from the same area as those identified by
Beier (1961)
, who added some characteristics with respect to the original description (1930). Differences found by
Callaini (1988)
involved the shape of the carapace and, especially, the chelal teeth, with those of the fixed finger teeth being distinctly longer than those on the movable finger, which are very flattened in the basal portion. Our observations confirm Callaini’s description, as opposed to Beier’s (
1961
,
1963
) in this respect. However, it should be noted that
Beier (1961)
compared
N. maroccanum
with
N. ischyrum
, the latter being a species known to have extremely flattened basal teeth on the movable chelal finger. Examination of the specimens from Xauen and Korifla does not reveal significant differences between them, except in the shape of the medial process of coxa I, which may be due to variability, and it can be concluded that they belong to the same species. In any event, it is clear that none of the populations assigned to
N. maroccanum
has a distinct large tooth on movable finger of chelicera (
Beier 1930
;
Callaini 1988
; this study), which provides an important difference from the new species described below.