Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae)
Author
Yasunaga, Tomohide
Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA;
Author
Tamada, Yui
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Hinami, Haruka
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Miyazaki, Ayana
Nagasaki West High School, Biology Club, Takenokubo 12 -
Author
Duwal, Ram Keshari
Visiting Researcher, Agriculture & Agri-Food Canada Environmental Health, K. W. Neatby: Bldg # 20, 960 Carling Avenue, Central Experimental Farm, Ottawa Ontario, Canada K
Author
Nagashima, Tetsuya
Nagasaki West High School, Biology Club, Takenokubo 12 -
text
Acta Entomologica Musei Nationalis Pragae
2019
Acta. Ent. Mus. Natl. Pragae
2019-02-26
59
1
71
99
http://dx.doi.org/10.2478/aemnp-2019-0007
journal article
5772
10.2478/aemnp-2019-0007
64a55ff4-0a71-491f-a4f4-9b2cd5a41431
1804-6487
4505468
027CE86F-9E75-44C3-A35E-E0C20BA4B693
Cleotomiris
Schuh, 1995
Diagnosis.
General shape rather ant-like (
Fig. 13
); small to moderate size (total length 3.0–
4.5 mm
); brown to fuscous basic coloration; more or less terete antennal segments II– IV; weakly swollen, trapezoidal pronotum that is not much constricted anteriorly (
Fig. 65
); ear-like ostiolar peritreme (
Figs 71
,
131
); presence of white fascia on clavus just posterior to scutellum (
Fig. 13
); possession of stridulatory device (currently confirmed in two species, see
Table 1
and
Figs 66–67
,
132–133
); weakly fleshy, apically convergent parempodia (
Fig. 68
); and short endosoma with an apical spine of various length and a developed secondary gonopore. Further diagnostic characters and detailed description were provided by
SCHUH (1984)
.
Distribution.
Known widely from the Sundaland west of the Wallacea, Indochina, southern
China
,
Philippines
and Japanese Okinawa Island (
YASUNAGA 2001
,
2012
).
Discussion.
Although this genus is superficially similar to
Cleotomiroides
, the male and female genitalic structures of these genera are significantly different from each other (see
YASUNAGA 2012
,
DUWAL et al. 2017
).
YASUNAGA (2012)
suggested that the similarity between
Cleotomiris
and
Cleotomiroides
was only superficial. Present observation also confirmed that some species of
Cleotomiroides
and
Wygomiris
are associated with broadleaf trees (on which immature forms were found, cf.
Figs 12, 17
) and lack the stridulatory mechanism. General shape of the female genitalia of
Cleotomiris
rather resembles that of
Wygomiris
. Almost all of the available specimens of these three genera were collected using UV light traps at well preserved subtropical or tropical forests. Therefore, members of these genera are currently considered to be arboreal.
Our examination by a SEM could find out the stridulatory devices in two
Cleotomiris
species,
C. miyamotoi
Yasunaga
(
Figs 66–67
) from
Okinawa
Island,
Japan
and an (presumably) undescribed species currently represented by a single female from
Yunnan Province
,
China
(NMPC). This unique Chinese species has rather developed FWS+MFP (
Figs 132–133
) which appear to retain the original function (stridulation) although it is superficially most similar to
C. miyamotoi
or
C. schneirlai
Schuh, 1984
(
Philippines
)
. From southern
China
(
Yunnan
,
Sichuan
) to northern Indochina, more than a few undetermined specimens representing at least three closely related species of
Cleotomiris
or
Cleotomiroides
are present. Therefore, definitive determination of these taxa is beyond a scope of the present study.