New Australasian Parathalassiinae (Diptera: Dolichopodidae sensu lato) Author Brooks, Scott E. 0000-0001-7915-1941 Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & scott. brooks @ agr. gc. ca; https: // orcid. org / 0000 - 0001 - 7915 - 1941 scott.brooks@agr.gc.ca Author Cumming, Jeffrey M. 0000-0002-3619-3700 Diptera Unit, Canadian National Collection of Insects, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, Ontario, K 1 A 0 C 6, CANADA & jeff. cumming @ agr. gc. ca; https: // orcid. org / 0000 - 0002 - 3619 - 3700 jeff.cumming@agr.gc.ca text Zootaxa 2022 2022-09-22 5188 6 521 543 journal article 148081 10.11646/zootaxa.5188.6.2 ecb5de27-f5fe-4b16-903b-633548186b0a 1175-5326 7103618 BF01B6B4-3415-41A1-86A0-F4B187541A55 Microphorella viticula sp. nov. ( Figs 1, 2 , 49–55 ) urn:lsid:zoobank.org:act: B4BCEB5C-323D-48AF-AE00-604896D7A5EC Type material. HOLOTYPE ♂ labelled: “ AUST : NSW: Blue Mtns. N.P./ Blackheath, Govetts/ Leap [ ca 33°38′S 150°19′E ], 4.iv.1994 / B.J. Sinclair / ex. dry scler. creek”; “ HOLOTYPE / Microphorella viticula / Brooks & Cumming [red label]” ( CNC ). Diagnosis. Males of M . viticula sp. nov. are distinguished from the other known Australasian Microphorella by the following combination of features: body and legs dark brown with black setation ( Fig. 49 ); palpus dark and narrowly ovate ( Fig. 51 ); hypopygium ( Figs 53–55 ) with phallus greatly elongated, slender and tightly curled; hypandrium relatively small, slightly wider than epandrial lamellae in lateral view. Description. Male ( Fig. 49 ): Body length 1.32 mm, wing length 1.35 mm. Similar to M. bungle sp. nov. except as follows: Head ( Figs 50, 51 ): 2 pairs of inclinate fronto-orbitals, posterior pair smaller; postocular setae slightly longer and pale brown; postpedicel 2X longer than wide, with basal half round and distal half narrow; arista-like stylus 1.8X length of postpedicel. Thorax ( Fig. 50 ). Legs ( Fig. 49 ). Wing ( Fig. 52 ). Abdomen ( Figs 49, 53–55 ): Hypopygium (some details not visible on unique undissected holotype ): Left ventral epandrial process longer. Ventral lobe of right surstylus apparently short, with a few apical setae. Hypandrium with posterior margin drawn into short pointed tip projecting dorsally and supporting protruding phallus. Ventral part of left postgonite lobe (medial to base of left ventral epandrial process) apparently developed into large flap-like lobe. Phallus greatly elongated, slender and tightly curled. Female. Unknown. Distribution. This species is known only from the type locality of Govetts Leap in Blue Mountains National Park, New South Wales , Australia ( Figs 1, 2 ). Etymology. From the Latin viticula , meaning tendril, in reference to the long thin curled phallus of the male. Remarks. The single holotype male is stated as having been taken in dry sclerophyl habitat near the creek ( Fig. 2 ), but was likely collected where the trail crosses the stream (B.J. Sinclair, pers. comm., 2022). Aside from the difference in phallic structure, the hypopygial morphology of M. viticula sp. nov. is very similar to M. bungle sp. nov. and the species appear closely related.