New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae) Author Perkins, Philip D. text Zootaxa 2011 2011-10-28 3074 1 198 journal article 31530 10.5281/zenodo.1050060 2d28bebf-e3c5-457b-9152-8a6877ded281 1175-5326 1050060 Ampla Complex Three species comprise the Ampla Complex: H. ampla , H. manabica , and H. cochabamba . These are medium sized to large species (ca. 1.44–1.97 mm ), which have a wide body form, are uniformly darkly colored dorsally, and lack a pronotal scintilla ( Figs. 126–127 , 130 ). The mesoventral intercoxal process (P2) is comparatively narrow, and the metaventrite does not have plaques. The male genitalia in this complex have a basic plan similarity, having the apex with three processes (one from the main piece and two from the distal piece), the gonopore bearing process is tube-like, pointing from the right to the left side, and the parameres are relatively short and originate near the distal piece ( Figs. 128–129 , 132 ). Species in this complex are currently known from Bolivia ( H. cochabamba ) and Ecuador ( H. ampla and H. manabica ).