<document id="84E2B5D64F1C74F863B28D6EED2E50A7" ID-CLB-Dataset="3574" ID-DOI="10.11646/zootaxa.5067.3.3" ID-GBIF-Dataset="937e9cfe-ea1e-4891-b147-91ab13a31d78" ID-ISSN="1175-5326" ID-Zenodo-Dep="5681919" ID-ZooBank="F64E5226-B4DD-44A3-A83E-E05928B82F4B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636617232589" checkinUser="plazi" docAuthor="Diago-Toro, María F., García-Cobos, Daniela, Brigante-Luna, Giovanni D. &amp; Vásquez-Restrepo, Juan D." docDate="2021" docId="03C787E7C317FF9EFF419AC25B454219" docLanguage="en" docName="zootaxa.5067.3.3.pdf" docOrigin="Zootaxa 5067 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loxopholis caparensis" docType="treatment" docVersion="8" lastPageNumber="389" masterDocId="FFFEFF9FC31BFF92FFD69B3C5E13410C" masterDocTitle="Fantastic lizards and where to find them: cis-Andean microteiids (Squamata: Alopoglossidae &amp; Gymnophthalmidae) from the Colombian Orinoquia and Amazonia" masterLastPageNumber="400" masterPageNumber="377" pageNumber="389" updateTime="1699468217192" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="916E1C9C132B3DD9283666F25D9E0F63">Fantastic lizards and where to find them: cis-Andean microteiids (Squamata: Alopoglossidae &amp; Gymnophthalmidae) from the Colombian Orinoquia and Amazonia</mods:title>
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<mods:namePart id="00E81F89D4BF084082F0F99FA4AF2A5B">Diago-Toro, María F.</mods:namePart>
<mods:affiliation id="C4A0ACB49C923F5474D6B2F609960907">Programa de Biología, Facultad de Ciencias Básicas y Tecnologías, Universidad del Quindío, Armenia, Quindío, Colombia.</mods:affiliation>
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<mods:namePart id="49E60F2644DE9D47A2F32A62BCF0A365">García-Cobos, Daniela</mods:namePart>
<mods:affiliation id="8BCCD44C5F4037DEBBFE7F92D5117655">Subdirección de Investigaciones, Colecciones Biológicas, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, carrera 8 # 15 - 08, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia.</mods:affiliation>
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<mods:namePart id="04830A15AC48AD86FC9B0CCFA1D5347D">Brigante-Luna, Giovanni D.</mods:namePart>
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<mods:affiliation id="2A3B4C3F800B934D20D549FD4FC11678">Grupo de investigación de Biodiversidad del Caribe Colombiano, Semillero de Herpetología, Programa de Biología, Universidad del Atlántico. https: // orcid. org / 0000 - 0001 - 6823 - 2383</mods:affiliation>
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<mods:namePart id="919A282567D58CBA02101BAADAAB24F2">Vásquez-Restrepo, Juan D.</mods:namePart>
<mods:affiliation id="C3B7EB6ABF307BA6786039ABCB1DEED4">Programa de Posgrado en Ciencias Biológicas, Instituto de Geología, Universidad Nacional Autónoma de México, Ciudad de México, México.</mods:affiliation>
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<treatment id="03C787E7C317FF9EFF419AC25B454219" ID-DOI="http://doi.org/10.5281/zenodo.5699697" ID-GBIF-Taxon="190305920" ID-Zenodo-Dep="5699697" LSID="urn:lsid:plazi:treatment:03C787E7C317FF9EFF419AC25B454219" httpUri="http://treatment.plazi.org/id/03C787E7C317FF9EFF419AC25B454219" lastPageNumber="389" pageId="12" pageNumber="389">
<subSubSection id="C374657AC317FF9EFF419AC25C654314" box="[151,630,510,537]" pageId="12" pageNumber="389" type="nomenclature">
<paragraph id="8BD136F1C317FF9EFF419AC25C654314" blockId="12.[151,630,510,537]" box="[151,630,510,537]" pageId="12" pageNumber="389">
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<taxonomicName id="4C6E4D72C317FF9EFF419AC25C654314" authority="(Esqueda, 2005)" baseAuthorityName="Esqueda" baseAuthorityYear="2005" box="[151,630,510,537]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="caparensis">
<emphasis id="B91AEAE3C317FF9EFF419AC25C654314" bold="true" box="[151,630,510,537]" pageId="12" pageNumber="389">
<emphasis id="B91AEAE3C317FF9EFF419AC25FB44314" bold="true" box="[151,423,510,537]" italics="true" pageId="12" pageNumber="389">Loxopholis caparensis</emphasis>
(
<bibRefCitation id="EFFF4B00C317FF9EFE6E9AC25C7D4314" author="Esqueda, L. F." box="[440,622,510,536]" pageId="12" pageNumber="389" pagination="33 - 42" refId="ref16263" refString="Esqueda, L. F. (2005) Un nuevo Leposoma (Squamata: Gymnophthalmidae) de la Reserva Forestal de Caparo, estado Barinas, Venezuela. Herpetotropicos, 2 (1), 33 - 42." type="journal article" year="2005">Esqueda, 2005</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374657AC317FF9EFF41997A5B454219" pageId="12" pageNumber="389" type="distribution">
<paragraph id="8BD136F1C317FF9EFF41997A5B834388" blockId="12.[151,1437,582,789]" pageId="12" pageNumber="389">
<emphasis id="B91AEAE3C317FF9EFF41997A5F20436C" bold="true" box="[151,307,582,608]" pageId="12" pageNumber="389">Distribution:</emphasis>
Known only from the Venezuelan and Colombian plains, in the Orinoco River basin (
<bibRefCitation id="EFFF4B00C317FF9EFB23997A5B84436D" author="Esqueda, L. F." box="[1269,1431,582,609]" pageId="12" pageNumber="389" pagination="33 - 42" refId="ref16263" refString="Esqueda, L. F. (2005) Un nuevo Leposoma (Squamata: Gymnophthalmidae) de la Reserva Forestal de Caparo, estado Barinas, Venezuela. Herpetotropicos, 2 (1), 33 - 42." type="journal article" year="2005">Esqueda 2005</bibRefCitation>
; 
<bibRefCitation id="EFFF4B00C317FF9EFF4199565FCD4389" author="Sanchez-Pacheco, S. J. &amp; Rueda-Almonacid, J. V. &amp; Caicedo-Portilla, J. R. &amp; Souza, S. M." box="[151,478,618,645]" pageId="12" pageNumber="389" pagination="53 - 57" refId="ref18997" refString="Sanchez-Pacheco, S. J., Rueda-Almonacid, J. V., Caicedo-Portilla, J. R. &amp; Souza, S. M. (2016) First record of Leposoma caparensis from Colombia, with confirmation for the presence of Ptychoglossus myersi and P. plicatus (Squamata: Gymnophthalmidae). Salamandra, 52 (1), 53 - 57." type="journal article" year="2016">
Sánchez-Pacheco 
<emphasis id="B91AEAE3C317FF9EFEB499575F894388" box="[354,410,618,644]" italics="true" pageId="12" pageNumber="389">et al.</emphasis>
2016
</bibRefCitation>
). In 
<collectingCountry id="F3797661C317FF9EFDD999565C6E4388" box="[527,637,618,644]" name="Colombia" pageId="12" pageNumber="389">Colombia</collectingCountry>
from the 
<collectingRegion id="49AAF813C317FF9EFD3F99565DAC4388" box="[745,959,618,644]" country="Colombia" name="Arauca" pageId="12" pageNumber="389">Arauca department</collectingRegion>
in the northernmost Orinoquia (
<figureCitation id="13552A74C317FF9EFAFF99565B974388" box="[1321,1412,618,644]" captionStart="FIGURE 4" captionStartId="4.[151,250,1888,1912]" captionTargetBox="[217,1370,181,1864]" captionTargetId="figure-22@4.[217,1370,181,1864]" captionTargetPageId="4" captionText="FIGURE 4. Distribution maps for the cis-Andean microteiid lizards in Colombia (Alopoglossidae + Gymnophthalmidae). Circles are examined specimens, while squares are records from literature (see Supplementary Files for specific references). Red arrows indicate new records for Colombia. Black arrows show a zoom in to regions where several records overlap. The light green shadow indicates the Orinoco River basin, and the dark green the Amazon River basin." figureDoi="http://doi.org/10.5281/zenodo.5681929" httpUri="https://zenodo.org/record/5681929/files/figure.png" pageId="12" pageNumber="389">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BD136F1C317FF9EFF1199B25B454219" blockId="12.[151,1437,582,789]" pageId="12" pageNumber="389">
<emphasis id="B91AEAE3C317FF9EFF1199B25F9243A4" bold="true" box="[199,385,654,680]" pageId="12" pageNumber="389">Similar species:</emphasis>
Currently, 
<taxonomicName id="4C6E4D72C317FF9EFE2A99B25CE443A4" baseAuthorityName="Esqueda" baseAuthorityYear="2005" box="[508,759,654,680]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="caparensis">
<emphasis id="B91AEAE3C317FF9EFE2A99B25CE443A4" box="[508,759,654,680]" italics="true" pageId="12" pageNumber="389">Loxopholis caparensis</emphasis>
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seems to have an allopatric distribution from other cis-Andean species of the genus. However, it can be differentiated from all the other eastern species of 
<taxonomicName id="4C6E4D72C317FF9EFB7E998E5B3743C0" box="[1192,1316,690,716]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="genus">
<emphasis id="B91AEAE3C317FF9EFB7E998E5B3743C0" box="[1192,1316,690,716]" italics="true" pageId="12" pageNumber="389">Loxopholis</emphasis>
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in 
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in having smooth or weakly keeled ventral scales (keeled in 
<emphasis id="B91AEAE3C317FF9EFCB799EB5AB843FC" box="[865,1195,726,752]" italics="true" pageId="12" pageNumber="389">
<taxonomicName id="4C6E4D72C317FF9EFCB799EB5DFF43FC" baseAuthorityName="Cope" baseAuthorityYear="1886" box="[865,1004,726,752]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="parietalis">L. parietalis</taxonomicName>
, 
<taxonomicName id="4C6E4D72C317FF9EFC2199EB5AB843FC" baseAuthorityName="Muller" baseAuthorityYear="1923" box="[1015,1195,727,752]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="percarinatum">L. percarinatum</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C6E4D72C317FF9EFB3E99EB5B9843FC" baseAuthorityName="Avila-Pires" baseAuthorityYear="1995" box="[1256,1419,726,752]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="snethlageae">
<emphasis id="B91AEAE3C317FF9EFB3E99EB5B9843FC" box="[1256,1419,726,752]" italics="true" pageId="12" pageNumber="389">L. snethlageae</emphasis>
</taxonomicName>
); and rhomboid dorsal scales (hexagonal in 
<taxonomicName id="4C6E4D72C317FF9EFDA699C75CEB4218" baseAuthorityName="Ayala &amp; Harris" baseAuthorityYear="1982" box="[624,760,762,788]" class="Reptilia" family="Gymnophthalmidae" genus="Loxopholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="389" phylum="Chordata" rank="species" species="hexalepis">
<emphasis id="B91AEAE3C317FF9EFDA699C75CEB4218" box="[624,760,762,788]" italics="true" pageId="12" pageNumber="389">L. hexalepis</emphasis>
</taxonomicName>
). For a more detailed description see 
<bibRefCitation id="EFFF4B00C317FF9EFB4899C65B414219" author="Esqueda, L. F." box="[1182,1362,762,789]" pageId="12" pageNumber="389" pagination="33 - 42" refId="ref16263" refString="Esqueda, L. F. (2005) Un nuevo Leposoma (Squamata: Gymnophthalmidae) de la Reserva Forestal de Caparo, estado Barinas, Venezuela. Herpetotropicos, 2 (1), 33 - 42." type="journal article" year="2005">Esqueda (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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