Palaeoparasycorax globosus Stebner et al., 2015 from the mid-Cretaceous amber of Myanmar: Description of a mating pair Author Azar, Dany State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Lebanese University, Faculty of Science II, Natural Sciences Department, Fanar - El-Matn, PO box 26110217, Lebanon Author Huang, Di-Ying State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China text Zootaxa 2023 2023-12-29 5396 1 94 104 https://www.mapress.com/zt/article/download/zootaxa.5396.1.14/52607 journal article 283447 10.11646/zootaxa.5396.1.14 5f91b413-59f8-4640-8df2-b6c051157350 1175-5326 10441131 10BDE7AD-B433-41A8-92C3-02B3F503CA20 Palaeoparasycorax globosus Stebner et al ., 2015 Figs 1‒4 Type species. Palaeoparasycorax globosus Stebner et al ., 2015 . Other species. Palaeoparasycorax suppus Stebner et al ., 2015 . Material. Specimen NIGP 203548 A (male) and B (female). FIGURE 1. Palaeoparasycorax globosus Stebner et al. , 2015 , specimen number NIGP203548. A , B , Habitus of mating pair. C , Head of male. D , Antenna of male. Scales bars = 0.5 mm in A , B , 0.1 mm in C , D . Amended diagnosis. Antenna with 14 flagellomeres, the apical one (apiculus) clearly distinct and half-length of the preapical one, ascoids not discernable, terminal palpus segment longest, sclerotized and distally broader than at its base; wing with Sc short and terminating in acute angle in R 1, sc-c crossvein present; r4+5-m crossvein absent; number of visible abdominal segments reduced ( i.e. , less than 8); male terminalia inverted; gonostylus upright, with distal portion blunt with one terminal and two subterminal long and strong spiniform setae; gonocoxite elongate, very slightly shorter than gonostylus; epandrial lobe well developed and ovaliform; female terminalia with reduced pointed cercus, slightly directed downward; 8 th sternite well developed, concave in lateral view, with sharp pointed apex ended by an apical strong seta, upward directed and surpassing cercus level. FIGURE 2. Palaeoparasycorax globosus Stebner et al. , 2015 , specimen number NIGP203548, wing of male. A , Microphotograph with compound microscope. B , Microphotograph with fluorescent compound microscope. C , Line drawing. Scale bars = 0.3 mm. Locality and horizon. Hukawng Valley of the northern state of Kachin ; latest Albian to earliest Cenomanian . Description of male. Body length without head and without gonostyli: 1.05 mm (same length as the holotype ). Head . With oval eyes ( Fig. 1C ); mouthparts short, non-functional, palpus ( Fig. 1C ) 4-segmented, terminal segment slightly sclerotized increasing in width distally; antennae longer than wing, with 14 flagellomeres ( Fig. 1D ), the apical one (apiculus) clearly distinct and half-length of the preapical one, first flagellomere longest, almost double in length than the two successive ones; ascoids not discernable; scape short, pedicel subspherical, with a short but evident apical neck. Thorax . Mesonotum setose; legs elongate without distinctive features; wing ( Fig. 2 ) 1.15 mm long, 0.49 mm large, with R 4+5 ending at the broad wing apex; Sc short, ending in a sharp angle in radial vein ( 0.4 mm from wing base), with a long and faint sc-c crossvein; radius with 4 veins, R4+5 fused; radial fork slightly basal to medial one; CuP [A] vein short almost not reaching wing margin. FIGURE 3. Palaeoparasycorax globosus Stebner et al. , 2015 , specimen number NIGP203548, male and female terminalia in mating. A , Microphotograph with compound microscope. B , Microphotograph with fluorescent compound microscope. C , Microphotograph with confocal microscope with color-coded projection. D , Microphotograph with confocal microscope. Scale bars = 0.1 mm. Abdomen . 0.5 mm long, with six visible segments; terminalia ( Figs 3 , 4B, 4C ) of inverted type , gonostylus upright, with distal portion blunt with one terminal and two subterminal long and strong spiniform setae; gonocoxite elongate, very slightly shorter than gonostylus; epandrial lobe well developed and ovaliform. Description of female. Body length without head: 1.5 mm . Head . Badly preserved with oval eyes; mouthparts relatively short but longer than those of the male, palpus 4- segmented, terminal segment slightly sclerotized increasing in width distally as in male; antenna (only one preserved) longer than wing, with 14 flagellomeres, the apical one (apiculus) clearly distinct and half-length of the preapical one, first flagellomere longest, almost double in length than the two successive ones; ascoids not discernable. Thorax . Mesonotum setose; legs elongate without distinctive features; wing with same venation as male but larger. Abdomen . 1.12 mm long; terminalia ( Figs 3 , 4A, 4C ) with reduced pointed cercus, slightly directed downward; 8 th sternite well developed, concave in lateral view, with sharp pointed apex ended by an apical strong seta, upward directed and surpassing cercus level. FIGURE 4. Palaeoparasycorax globosus Stebner et al. , 2015 , specimen number NIGP203548. A , Line drawing of female terminalia. B , Line drawing of male terminalia. C , Line drawing of male and female terminalia in copulation. Abbreviations: ae = aedeagus; ce = cercus; ep lb = epandrial lobe; goncx = gonocoxite; gonst = gonostylus; pm = paramere; 8th st = 8 th sternite. Scale bars = 0.1 mm. Remarks. Palaeoparasycorax globosus differs from all remaining Sycoracinae by the distinctive absence of r4+5-m crossvein. Male smaller and thinner than female; female 1.5x longer than male and 1.8x wider ( Fig. 1A, B ). Copulation with end-to-end opposite position during courtship ( Fig. 1A, B ). The male captures the female terminalia with its gonostyli and penetrates the two filaments of the aedeagus attached basally to the parameres, while the female upwards the 8 th sternite to enable mating.