Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae) Author Selnekovič, Dávid D930D757-D929-4248-A29C-A857E0505345 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. david.selnekovic@uniba.sk Author Jäch, Manfred A. D749707A-8823-4110-8D51-BF5AF4E2820F Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria. manfred.jaech@nhm-wien.ac.at Author Kodada, Ján 6E88BFBB-8769-44EC-8285-29E357CEC064 Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia. jan.kodada@uniba.sk text European Journal of Taxonomy 2024 2024-09-11 957 1 229 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247 journal article 10.5852/ejt.2024.957.2651 2118-9773 13773385 9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3 Elmomorphus striatellus Delève, 1968 Figs 6–7 , 12A , 107A , 108A Elmomorphus striatellus Delève, 1968: 150–151 (original description; figure). Differential diagnosis Elmomorphus striatellus ( Fig. 6 ) is characterised by having the dorsal surface covered with plastron, except for a narrow area along the posterior pronotal margin. The presence of plastron on the prosternal process is also a characteristic feature. Each elytron possesses nine shallow longitudinal striae. Phallobase and parameres are rather short and robust ( Fig. 7 ). Proximal and distal part of bursa copulatrix with sclerotised spines ( Fig. 12A ). This species most closely resembles E. siamensis sp. nov. , but it differs from the latter in the presence of elytral striae and the larger size, TL: ♂♂ 3.36–3.84 mm (3.53 ± 0.16, n= 10), ♀♀ 3.41–3.77 mm (3.54 ± 0.12, n =10) versus TL: ♂♂ 2.73–3.20 mm (3.00± 0.16, n= 10), ♀♀ 2.78–3.25 mm (2.99 ± 0.14, n= 10) in E. siamensis ; the parameres are distinctly broader, with broadly rounded apices; the phallobase is shorter and wider ( Fig. 7 ); the bursa copulatrix has distinctly fewer spines ( Fig. 12A ). Type material Holotype VIETNAM • ♂ ; “ ♂ | VIETNAM , Prov. Nghe-An, forestière Quy-shâu, à la lumière | forêt pluv. trop. semidecid., 24.VIII.1963 , T. PÓCS | Holotypus 1967 Elmomorphus striatellus Delève [white label with red margins] | Holotype [red label] | J. Delève det. 1967 Elmomorphus striatellus n. sp. | Elmomorphus striatellus Deleve ”; HNHM . Paratype VIETNAM • 1 ♀ ; same collection data as for holotype; “ ♀ ”; “ Paratypus 1967 Elmomorphus striatellus Delève ”; HNHM . Additional material examined LAOS – Nam Ngang Province • 2 ♂♂ , 4 ♀♀ ; “LAOS-CE, 1-18.v.2001 , Boli Kham Xai prov. , 18°21′N 105°08′E BAN NAPE ( 8 km NE), ~ 600m , C.L.Peša leg.”; CKB , NMW . VIETNAM – Hoa Binh Province • 1 ex. ; “ Vietnam , Hoa Binh , 4.-7. 6. 1986 Ha Son Binh prov. [today: Hoa Binh Province ] lgt. J. Rybníček”; CKB . – Lao Cai Province • 8 ♂♂ , 3 ♀♀ , 6 ex. ; “N-VIETNAM 1991 Sapa ( Lao Cai ) leg. E. Jendek | 26.5.-10.6., 22°20′N 103°50′E ”; CKB , NMW • 2 ex. : “N VIETNAM 5. 1991 SA PA, 1200 m Hong Lien Sond D. T. TU leg.”; CKB , NMW . – Ninh Binh Province • 5 ex. ; “N-VIETNAM 1991 Cuc-Phuong Nationalpark | 2.-12.V. 400 km S Hanoi leg. Jendek ”; CKB , NMW • 2 ♀♀ ; “ VIETNAM Cuc Phuong 2.-11.V.91 Strnad Jan lgt.”; CKB • 1 ex. ; “N-VIETNAM, Cuc Phuong NP, N 20° 17.572′ E 105° 40.052′, 270m , 22.5. – 24.5.2015 , leg. A. Skale ”; NME • 1 ex. ; “N-VIETNAM, P: Nin Binh 90 Km SW Hanoi Cuc Phuong NP env. centre 07.V. 2017 , 270 m, 21°17′572″N, 105°40′052″E , leg. A. Weigel LFF [ light trap ] primary forest”; CWW . – Thua Thien Hue Province • 22 ex. ; “ VIETNAM , Thua Thien Hue Pr., Phong Điền Distr., Phong Mỹ, 16°31′15.3″N , 107°15′00.3″E , 27 m a.s.l. , 18.11.2022 , Selnekovič & Kodada leg. (15)”; CKB . – Vinh Phúc Province • 3 ♀♀ ; “NORTH VIETNAM Pr. VINH PHU[C], TAM DAO 5.- 25.5.1990 O. ŠAUŠA LGT.”; CKB • 2 ex. ; “N-VIETNAM: Thai Ngayen Prov. [ Vinh Phúc Province ], vic.[inity of] Ngoc Thanh , Me Linh ( IEBR [ Institute of Ecology and Biological Resources ] station) 21°23′3″N 105°42′44″E 60-80m , 12.V.2012 leg. A. Skale ”; NME • 6 ex. ; “N- VIETNAM , P: Tai Nguyen [ Vinh Phúc Province ], vic. Ngoc Than , Me Linh ( IEBR station) 13.V.2017 , 21°23′3″N , 105°42′44″E , 65 m NN leg. A. Weigel LFF”; CWW . – Yen Bai Province • 1 ♂ , 2 ♀♀ ; “ VIETNAM NORTH, YEN BAI , 6.-25. 5. 1990 O ŠAUŠA LGT.”; CKB • 2 ex. ; “N VIETNAM (Tonkin) pr. Hoang Lien Son YEN BAI 10.V.1990 P. Pacholátko leg.”; CKB , NMW . – Thua Thien Hue Province • 9 ex. ; “C-VIETNAM, Th ừa Thiên-Huế Pr. ca. 30km W Hu ế, River valley 16°17′59″N , 107°20′22″E , 80m 10.V.2019 , leg. A. Skale , LF [ light trap ]”; NME • 4 ex. ; “C-VIETNAM, Thua Thien-Hue Prov. ca. 30 km W Hue , 80 m , 16°17′59″N 107°20′22″E , 10.V.2019 , by light river valley leg. A. Weigel ”; CWW • 2 ex. ; same collection data as for preceding, but different label: “C- VIETNAM , Thua Thien-Hue , ca. 30 km W Hue , River valley , 80 m , LF [ light trap ] 16°17′59″N 107°20′22″E , 10.V.2019 , leg. A. Weigel ”; CWW • 6 ex. ; “C-VIETNAM, Th ừa Thiên-Huế Prov. , ca. 30km NW Hu ế, Sôn Ô Lâu River , 16°31′3″N , [1] 07°15′36″E , 30m , 11.V.2019 , leg. A. Skale , LF [ light trap ]”; NME • 4 ex. ; “C- VIETNAM , Thua ThienHue ca. 30km NW Hue Son O Lau River valley , 30 m , by light 16°31′3″N 107°15′36″E , 11.V.2019 , leg. A. Weigel ”; CWW . Fig. 6. Elmomorphus striatellus Delève, 1968 , male from Ban Nape, Laos (CKB), TL: 3.50 mm. Fig. 7. Elmomorphus striatellus Delève, 1968 , aedeagus, specimens from Sa Pa, Vietnam (CKB). A . Lateral aspect. B . Lateral aspect. C . Ventral aspect. Scale bar: 0.1 mm. Type locality Vietnam , Nghệ An Province, Quỳ Châu. Redescription Measurements (mm): TL: ♂♂ 3.36–3.84 (3.53 ± 0.16, n =10), ♀♀ 3.41–3.77 (3.54 ± 0.12, n=10); PL: ♂♂ 0.70–0.85 (0.78 ± 0.04, n =10), ♀♀ 0.72–0.85 (0.77± 0.04, n=10); PW: ♂♂ 1.33–1.50 (1.39 ± 0.05, n =10), ♀♀ 1.37–1.47 (1.41 ±0.03, n =10); EL: ♂♂ 2.60–2.99 (2.75± 0.12, n =10), ♀♀ 2.70–2.93 (2.77 ±0.08, n=10); EW: ♂♂ 1.56–1.76 (1.66± 0.07, n= 10), ♀♀ 1.63–1.79 (1.69± 0.05, n= 10); PhL: 0.53–0.62 (0.59 ± 0.03, n=10); PrL: 0.35–0.40 (0.38 ± 0.02, n=10). Body ovate ( Fig. 6 ), widest behind midlength of elytra, strongly convex dorsally, highest point in anterior third of elytra. Colouration black, except for reddish brown mouthparts, trochanters and tarsi. Pubescence on dorsal surface consists of short thin decumbent yellowish setae arising from small round setiferous punctures. Plastron present on entire cranial surface, on pronotum, except for narrow transverse area before scutellum, and on entire elytral surface. Plastron present on entire ventral surface, except for median keel of prosternal process and median part of metaventrite. Cranial surface entirely covered with plastron; round setiferous punctures separated by ca 0.5× puncture diameter, each puncture ca 0.75 × as wide as an eye facet. Labrum transverse, anterior margin straight, anterolateral angles rounded; exposed part microreticulate, with small setiferous punctures; setae short, decumbent, similar in both sexes. Clypeus with straight anterior margin, with a row of short thin setae. Eyes large, oval, moderately protruding; ID: ♂♂ 0.42–0.49 mm (0.45± 0.02, n= 10), ♀♀ 0.43–0.48 mm (0.45 ± 0.02, n=10). Antennae short, 11-segmented, densely covered with setae. Pronotum transverse, widest at base, moderately convex, PW/PL: ♂♂ 1.69–1.93 (1.84± 0.07, n=10), ♀♀ 1.69–1.93 (1.84 ± 0.07, n= 10); surface entirely covered with plastron, except for narrow transverse area before scutellum; round setiferous punctures as wide as those on cranium, separated by ca 1–2 × puncture diameters; anterolateral pronotal angles moderately deflexed, produced and acute; lateral sides weakly curved in the middle. Prosternal process covered with plastron, except on median keel; lateral and posterior margins rounded; lateral parts raised, without clusters of long setae in males; median keel strongly arcuate. Scutellum longer than wide, smooth, with several setiferous punctures. Metaventrite covered with plastron, except for narrow, posteriorly narrowed median part; disc flat, without clusters of long setae in males; lateral sides of metaventral process raised. Elytra ovate, convex, widest around middle, lateral sides regularly rounded; EL/EW: ♂♂ 1.58–1.71 (1.66± 0.05, n =10), ♀♀ 1.60–1.68 (1.64 ± 0.03, n=10); surface entirely covered with plastron; each elytron with nine shallow striae; small, round setiferous punctures scattered over entire surface. Tibiae straight; male protibia slightly longer than tarsus; PTL/PL: ♂♂ 1.06–1.23 (1.15± 0.05, n =10), ♀♀ 1.09–1.34 (1.20 ±0.06, n=10). Terminal tarsomere in male foreleg ca 1.25 × as long as all preceding segments combined; foreclaws similar in both sexes, narrow and strongly curved. Ventrites entirely covered with plastron. Lateral margins of intercoxal process flat; admedian keels absent. Ventrite 5 evenly convex, apex narrowly truncate in both sexes; females with indistinct longitudinal keel at apex. Aedeagus ( Fig. 7 ): phallobase short and robust, PhL/PrL: 1.38–1.67 (1.55 ±0.10, n=10); parameres wide in lateral aspect, apices broadly rounded; penis long, almost reaching apex of parameres, apex narrowly rounded; sclerotised fibula long and slender. Proximal and distal part of bursa copulatrix with few large spines ( Fig. 12A ). Secondary sexual dimorphism Average body dimensions similar in both sexes. Females can be distinguished by the presence of a small, hardly discernible longitudinal subapical keel on ventrite 5. Distribution Laos (first record), Vietnam ( Fig. 108A ).