A Revision of the Spider Genus Zorocrates Simon (Araneae, Zorocratidae)
Author
PLATNICK, NORMAN I.
Author
UBICK, DARRELL
text
American Museum Novitates
2007
2007-06-28
3579
1
44
http://www.bioone.org/doi/abs/10.1206/0003-0082%282007%293579%5B1%3AAROTSG%5D2.0.CO%3B2
journal article
10.1206/0003-0082(2007)3579[1:AROTSG]2.0.CO;2
0003-0082
5388198
Zorocrates fuscus
Simon
figures 1–5
; map 1
Zorocrates fusca
Simon, 1888: 212
(
two female
syntypes
from
Guanajuato
,
Mexico
, in MNHN, examined).
Zorocrates fuscus
:
Simon, 1892: 230
(emendation of gender).
Zorocrates
cf.
guerrerensis
:
Silva, 2003: 41
,
fig. 13a.
NOTE: In the original description, the type locality is presented only as ‘‘Mexique’’ but Simon later (1895b: 106) specified ‘‘Mexique central’’ and that later restriction corroborates the current label with the
syntypes
, which indicates that the specimens were collected in
Guanajuato
,
Mexico
, by
Dugès
.
DIAGNOSIS: Males resembles those of
Z. unicolor
but have a longer, fingerlike (rather than ledge-shaped) proximal lobe on the retrolateral tibial apophysis, a distally rounded median apophysis (
figs. 2, 3
), and a ventrally shorter tegular apophysis; the median apophysis does have a small, subdistal denticle, but it is directed dorsally and is therefore easy to overlook in ventral view (the denticle is more prolaterally directed and conspicuous in males of
Z. unicolor
). Females also resemble those of
Z. unicolor
but have deeper epigynal pockets with smaller, more medially situated openings connected to anteriorly expanded margins that parallel a longitudinal ridge between the pocket and the epigynal midpiece (
fig. 4
).
Figs. 1–5.
Zorocrates fuscus
Simon.
1.
Left male palp, prolateral view.
2.
Same, ventral view.
3.
Same, retrolateral view.
4.
Epigynum, ventral view.
5.
Same, dorsal view.
MALE: Total length 14. Carapace light brown with dusky triangular markings radiating from thoracic groove; abdomen gray, with lighter cardiac spot, posterior one-fifth with chevron pattern; femora light brown, more distal leg segments grading to brown on metatarsi and tarsi. Leg spination: femora: I r1-1-2; II p0-1-2, r1-1-2; III p1-1-1; IV r0-0-2; patellae IV r0-1-0; tibiae III r1-0-1; metatarsi I p1-1-1, v2-2-0, r1-1-0. Cribellum present. Retrolateral tibial apophysis short, with fingerlike proximal lobe; median apophysis hook-shaped, distally rounded; tegular apophysis sharply pointed (
figs. 1–3
).
Map 1. Records of the
Zorocrates fuscus
group (excluding
Z. unicolor
):
Z. fuscus
(dots),
Z. guerrerensis
(triangles; suspect record from
Panama
not plotted),
Z. gnaphosoides
(circles),
Z. ocampo
(open squares),
Z. mordax
(filled square).
FEMALE: Total length 16. Coloration as in male. Leg spination: femora: I, II p0-1-1, r1-1-0; IV p2-0-1, r0-0-1; patellae IV p0-0-0, r0-1-0; tibiae: I, II d0-0-0, p0-0-0, r0-0-0; III r1-0-1; IV r2-0-1; metatarsi: I p0-0-0, r0-0-0; II p0-0-0, r0- 0-1; III p1-1-2, r1-1-2; IV v3-2-2. Cribellum present. Epigynal midpiece triangular, with transverse anterior ledge; lateral pockets present, with relatively small openings; spermathecae with short anterior lobes (
figs. 4, 5
).
MATERIAL
EXAMINED:
MEXICO
:
Distrito Federal
:
no specific locality,
Sept. 1940
(
F. Bonet
, AMNH), 13,
Oct. 20, 1942
(
R. Ibarra
H., AMNH), 13,
Dec. 15, 1949
(AMNH), 13,
May 1955
(AMNH),
1♀
,
Mar. 1956
(UNAM), 13,
Sept. 2, 1958
(UNAM), 13,
Nov. 1958
(UNAM), 13; Azcapotzalco,
Oct. 27, 2002
(UNAM),
1♀
; Col. Medellín,
Oct. 31, 1941
(AMNH), 13;
Contreras, Aug.
9, 1942, elev.
2800 m
(C. Bolivar, AMNH),
1♀
,
Mar. 4, 1944
(M. Cardenas, AMNH),
1♀
;
Coyoacán
,
Sept. 1946
, elev.
2300 m
(
H. Wagner
, AMNH), 13,
Nov. 1, 2000
(AMNH), 23;
Cuauhtémoc
,
Mar. 2000
, in house (UNAM),
1♀
;
Mexico City
,
Oct. 1945
(H.
Wagner
, AMNH), 13, winter 1945–1946, in house (
H. Wagner
, AMNH), 13,
Dec. 25, 1975
, on building downtown (
V. Roth
, AMNH),
1♀
;
Miguel Hidalgo
,
Nov. 20, 2001
, on patio (UNAM), 13;
Pedregales
,
July–Aug. 1909
, under stone (
A. Petrunkevitch
, AMNH),
1♀
,
Petregal
[probably Pedregal],
Oct. 2, 1947
(
H. Wagner
, AMNH), 13.
Guanajuato
:
no specific locality (
Dugès
, MNHN AR230),
2♀
(
syntypes
).
Guerrero
:
summit,
4 mi
W Cacahuamilpa
,
18
°
41
9
N
,
99
°
34
9
W
,
Sept. 3, 1966
(
J., W. Ivie
, AMNH),
1♀
;
3 mi
N Chilpancingo
,
Nov. 18, 1946
(E.
Ross
, CAS),
1♀
;
72.6 mi
SE Ciudad Altamirano
, 2.1 mi
S Highway
51 on road to
Apaxtla
,
Nov. 11, 1987
, elev.
1520 m
(V.
Lee
, CAS),
1♀
;
Taxco
,
Oct. 1945
(
L. Isaacs
, AMNH), 13,
Apr. 1946
(
L. Isaacs
, AMNH), 13;
3 km
SE Tuxpan
,
Nov. 3, 1976
, elev.
1768 m
(E.
Ross
, CAS), 13,
2♀
.
Hidalgo
:
Apulco,
Oct.
6, 1947 (H.
Wagner
, AMNH),
2♀
;
El
Tablón,
7 mi
SE
Zimapán
,
20
°
40
9
N
,
99
°
20
9
W
,
Aug. 19, 1964
(gJ.,
W. Ivie
, AMNH), 33 (identified from scanning electron micrographs by
M. Ramírez
);
2 mi.
NE Jacala
,
Sept. 1, 1946
, elev.
1500 ft.
, pine-oak (C.
Bogert
, AMNH),
1♀
;
Pachuca
,
Jan. 28, 1943
(AMNH),
1♀
;
Parque Nacionale de El Chico
,
June 28, 1943
, elev.
3100 m
(C.
Bolivar
, F.
Bonet
,
Osorio
,
Pelaez
, AMNH),
1♀
;
Tasquillo
,
Tzindejeh
,
20
°
33
9
N
,
99
°
19
9
W
,
July 29, 1966
(
J., W. Ivie
, AMNH),
1♀
;
Tulancingo
, 1957 (UNAM), 33.
Jalisco
:
Jocotepec
,
July 31, 1965
(
Conant
, AMNH),
1♀
;
E end,
Lago de Chapala
,
Nov. 30, 1948
(H.
Leech
, CAS),
2♀
;
W side,
Laguna de Sayula
,
July 30, 1964
(W.
Gertsch
, J.
Woods
, AMNH),
2♀
; 22.7 mi S Puerto Vallarta,
Oct. 20, 1973
(S. Williams, C. Mullinex, CAS),
1♀
; 3.5 mi S Zocoalco,
Aug. 18, 1959
(C. Bogert, AMNH),
1♀
.
México
:
Aculco
,
Route
55 detour to
San Joaquín
,
Oct. 13, 2001
, disturbed pasture, elev.
2485 m
(
O. Francke
,
E. González
, UNAM), 13;
Malinalco
, ruin,
Oct. 25, 1973
(
S. Williams
,
C. Mullinex
, CAS),
2♀
;
Naucalpan
,
Nov. 27, 1997
, in house (
F. Alvarez
, UNAM), 13;
Reyes Iztacala
,
Oct. 25, 2001
, in house (C.
Durán
, AMNH), 13;
San Juan Teotihuacán
,
Nov. 4, 1939
(
C. Bogert
,
H. Vokes
, AMNH),
1♀
;
Tenancingo
,
Sept. 27–Oct. 7, 1946
, elev.
2050 m
(
H. Wagner
, AMNH), 23,
6♀
;
Tenango del Valle
,
Aug. 25–26, 1946
, elev.
2400 m
(
H. Wagner
, AMNH), 13;
Teotihuacán
ruins,
Aug. 13, 1977
, elev.
7400 ft
(C.
Griswold
,
T. Meikle
, CAS),
1♀
;
Tonatico
,
Dec. 1, 1996
(UNAM), 13.
Michoacán
:
Jiquilpan
,
19
°
59
9
N
,
102
°
41
9
W
,
May 9, 1963
(
W. Gertsch
,
W. Ivie
, AMNH),
3♀
;
Morelia
,
19
°
40
9
N
,
101
°
12
9
W
,
May 8, 1963
(
W. Gertsch
,
W. Ivie
, AMNH),
3♀
;
Pátzcuaro
,
Oct. 20–Nov. 13, 1955
(
C. Bogert
, AMNH),
1♀
;
24 mi
S Sahuayo on Highway
15 to
Morelia
,
Oct. 23, 1973
(
S. Williams
,
C. Mullinex
, CAS),
1♀
;
3 km
E Tuxpan
,
Jan. 6, 1989
, elev.
1825 m
(E.
Ross
, R.
Stecker
, CAS),
2♀
;
Zamora
,
Aug. 1, 1956
(
W. Gertsch
,
W. Ivie
, AMNH),
1♀
;
Zitácuaro
,
Dec. 6, 1943
, elev.
2000 m
(
C. Bolivar
, AMNH),
1♀
.
Morelos
:
Cuernavaca
,
Apr. 1959
(
N. Krauss
, AMNH),
1♀
; N Cuernavaca,
18
°
56
9
N
,
99
°
14
9
W
,
May 6, 1963
(W. Gertsch, W. Ivie, AMNH),
3♀
;
Oaxtepec
,
May 17, 1942
(
Correa
,
Cardenas
, AMNH),
1♀
;
Tepoztlán
,
Dec. 3, 1947
(UNAM),
1♀
.
Oaxaca
:
1.5 mi
S Carrizal
,
July 25, 1963
, elev.
9500 ft
(
G. Sluder
, AMNH),
1♀
;
El Punto
, road to
Ixtlán de Juárez
,
Aug. 19, 1961
(
C., M. Bogert
, AMNH),
2♀
;
5 mi
NE Mitla
, near ‘‘
El
Crucero’’ ruins,
Aug. 27, 1963
, on ridge, elev. ca.
6800–7200 ft
(M.
Bogert
,
G. Sluder
,
N. Bucknall
, AMNH),
1♀
;
Monte Alban
,
Dec. 11, 1943
(
E. Ross
, CAS),
1♀
;
0.5 mi E
Nochixtlán
,
1 mi
SE El Palmar
,
Dec. 11, 1948
(E.
Ross
, CAS),
1♀
;
9 mi
S Nochixtlán
,
17
°
20
9
N
,
97
°
12
9
W
,
May 1, 1963
(W.
Gertsch
,
W. Ivie
, AMNH),
1♀
.
Puebla
:
Acultzingo
,
July 4, 1963
(AMNH),
2♀
;
Ozumbilla
,
18
°
37
9
N
,
97
°
25
9
W
,
Apr. 25, 1963
(
W. Gertsch
,
W. Ivie
, AMNH),
1♀
;
Tehuacán
,
Nov. 8, 1939
(
C. Bogert
,
H. Vokes
, AMNH),
2♀
;
3 mi
NE
Zacatepec on Route
140,
June 30, 1963
(AMNH),
1♀
.
San Luis Potosı´
:
Route
70, 70 mi
W Valles
,
Feb. 19, 1970
, under roadside stone (
J. Cooke
, AMNH),
1♀
.
Tlaxcala
:
Tlaxcala
,
July 26, 1956
(
W. Gertsch
,
V. Roth
, AMNH),
1♀
.
Veracruz
:
Carrizal
,
Feb. 10, 1948
(
H. Wagner
, AMNH),
1♀
;
Jalapa
(BMNH 1905.4.28.1341- 1346),
1♀
;
Orizaba
(
Eislin
,
Vaslit
, MCZ 56234),
1♀
;
Perote
,
June 30, 1946
(
H. Wagner
, AMNH),
1♀
;
Zongolica
,
Mar. 1983
(AMNH),
1♀
.
Zacatecas
:
20 km
E Tlaltenango
,
Sept. 14, 1984
(W.
Pulawski
, CAS),
1♀
.
DISTRIBUTION: Central and southern
Mexico
(map 1);
one female
supposedly taken at
St. Ann’s
,
Trinidad
(
Aug. 13, 1958
, M.
Nieves
, AMNH) is presumed to be either mislabeled or introduced
.