The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae)
Author
Bosmans, Robert
Author
Kherbouche-Abrous, Ourida
Author
Benhalima, Souâd
Author
Hervé, Christophe
text
Zootaxa
2018
2018-07-30
4451
1
1
67
journal article
29293
10.11646/zootaxa.4451.1.1
62552028-dd96-4a85-bd83-5e86bd957be4
1175-5326
1445080
5DCE18B3-9DBA-48F5-AEF7-90A50B984EA5
Haplodrassus dentifer
Bosmans & Abrous
,
sp. n.
Figs 30‒40
,
49‒53
, map 2
Types
.
Holotype
♂
,
paratypes
1♂
2♀
from
Morocco
,
Souss-Massa
,
between Aoulouz and Tallliouine
(
N30°34’41”
W8°3’11”
),
600m
, stones in steppe,
4.II.1996
, J.
Van
Keer leg.; deposited in
RBINS
.
Etymology.
The species name is derived from the Latin noun
dens
(tooth) and the verb
ferre
(to wear) and refers to the presence of a tooth at the base of the embolus.
Diagnosis.
Haplodrassus dentifer
sp. n.
is closely related to
H. rufipes
. Males are at once recognized by the presence of a subterminal tooth at the prolateral margin of the embolic apophysis in
H. rufipes
(
Figs 131
,
142
), absent in
H. dentifer
(
Figs 35
,
51
), females by the converging, gradually narrowing antero-lateral branches of the areola (
Figs 40
,
52
), straight and truncate in
H. rufipes
(
Figs 132
,
143
).
Description.
Measurements: Male (n=5): Total length 5.6‒9,6 mm; carapace 1.22‒2.02 long, 0.99‒1.58 wide. Female (n=11): Total length
7.6‒9.6 mm
; carapace 1.96‒2.44 long, 2.44‒3.12 wide. Colour: Carapace yellowish brown with darkened eye region; chelicerae dark brown; sternum and legs yellowish brown; abdomen grey.
Male palp (
Figs 32‒36
,
49‒51
): Tibia 2.5 times as long as wide, gradually narrowing, slightly widened before the rounded tip (
Figs 33
,
50
); embolic apophysis nearly straight, without denticulations, with pointed tip; base of embolus with strong denticulate keel (
Figs 36
).
Epigyne (
Figs 38‒40
,
52‒53
): Anterior margin nearly as wide as areola; pro-fovea short, less than half the length of the fovea; foveal border marked by a strong medio-lateral tooth, but often obscured by the presence of concretions (compare
Figs 39 and 40
).
Further
material examined. AFRICA
:
ALGERIA
,
Laghouat
,
20 km
south of
Laghouat
(
N33°38’41”
E2°55’51”
),
740m
,
1♂
, pitfalls under
Zizyphus
bushes in a daya,
14.V.1990
,
R. Bosmans
leg. (
CRB
)
,
Ghardaia
:
Ksar
el
Atteuf
(
N32°28’30”
E3°44’57”
),
1♂
, pitfalls in alfalfa cultures,
25.III.2015
,
Y. Alioua
&
A. Hadj Mhammed
leg. (CYA)
,
Tamanrasset
,
Issek
(
N22°34’35”
E6°01’54”
),
1200m
,
1♀
,
II.1988
,
K. De Smet
leg. (
CRB
)
.
MOROCCO
,
Draâ-Tafilale
t
,
Col de Tagalm
,
SE Zebzat
,
1930m
,
2♀
, stone field,
19.IV.2012
, R.
Bosmans
&
J.
Van
Keer
leg. (
CJVK
,
CRB
)
,
Tinerhir
20 km
E. (
N31°30’36”
W5°31’48”
),
1350m
,
1♂
, litter in small palm yard,
6.II.1996
,
J.
Van
Keer
leg. (
CJVK
)
,
Tizi
‘n
Ikhsane
(
N30°26’56”
W7°31’24”
),
1650m
,
1♀
, stones in steppe,
6.II.1996
,
J.
Van
Keer
leg. (
CJVK
)
,
Fès-Meknès
, Missour (N33°2’56” W3°59’22”), 1♀,
7.XII.2001
, 1♀,
7.III.2002
, 2♀,
27.IV.2002
, pitfalls in steppe, R. Bosmans leg. (CRB),
Guelmim-Oued Noun
,
10 km
W.
Taghjijt
(
N29°04’02”
W9°17’52”
),
650m
,
1♀
, stones in steppe,
13.
II.2007
,
R. Bosmans
leg. (
CRB
)
,
Tata
(
N29°45’35”
N7°58’57”
E),
1♀
, litter in palm yards,
14.
II.2007
,
R. Bosmans
leg. (
CRB
)
,
Lâayoune-
Saguia
al Hamra
, Tarfaya
40 km
S., (
N27°35'53”
W12°57'
11'),
2♀
, stony desert,
11.
II.2007
, R.
Bosmans
&
J.
Van
Keer
leg. (
CRB
)
,
Marrakech-Safi
, Imi n’Tanoute (N31°10’16” W8°50’43”), 1♀, Coll. Simon 25299 (MNHN AR15687).
TUNISIA
,
Sousse
,
Hergla S.
(
N35°59’17”
W10°30’41”
),
10m
,
1♂
, pitfalls in salt marsh,
27.II‒8.III.2005
,
J. Van Keer
leg. (
CJVK
)
.
EUROPE
:
SPAIN
:
Cádiz:
Sierra
de los Pinos, Villalengua del
Rosario
(
N36°37’55”
W5°22’33”
),
1400m
,
1♀
, under stones in the crest of a limestone mountain,
16.II.2016
,
I. Sánchez
leg. (
CIS
)
.
FIGURES 30–40
.
Haplodrassus dentifer
sp. n.
(paratypes from Morocco, Aoulouz). 30. Male, dorsal aspect; 31. Female, dorsal aspect; 32. Cymbium of male palp, dorsal view; 33. Male palpal tibia, dorsolateral view; 34. Male left palp, retrolateral view; 35. Idem, ventral view; 36. Idem, retrolateral view; 37. Distal part of female palp, lateral view; 38‒39. Epigyne, ventral view, two different specimens; 40. Vulva, ventral view.
Ecology.
The specimens were collected in steppe vegetation, salt marshes, stony fields, dunes with scarce vegetation and palm yards, situated in the arid, semi-arid and desert climate zones of the Maghreb. Males and females were found from December to May.
Distribution.
This species occurs in
Algeria
,
Morocco
and
Tunisia
and was once collected in the South of
Spain
(map 2).