The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea Author Harzhauser, Mathias 0000-0002-4471-6655 Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at https: // orcid. org / 0000 - 0002 - 4471 - 6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard M. 0000-0002-7768-8494 Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com https: // orcid. org / 0000 - 0002 - 7768 - 8494 bernardmlandau@gmail.com Author Vermeij, Geerat J. 0000-0002-4471-6655 Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Geology, University of California at Davis, One Shields Avenue, Davis Ca 95616, USA. gjvermeij @ ucdavis. edu; https: // orcid. org / 0000 - 0002 - 6450 - 5687 & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at https: // orcid. org / 0000 - 0002 - 4471 - 6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2024 2024-06-14 5470 1 1 92 http://dx.doi.org/10.11646/zootaxa.5470.1.1 journal article 10.11646/zootaxa.5470.1.1 1175-5334 12189689 6A4270C2-D3F9-404F-91E7-4A73F2A99AE4 Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890) Figs 5A–C * Fasciolaria pyrulaeformis nov. form.—Hoernes & Auinger 1890: 265, pl. 32, figs 5–6. Fasciolaria ( Pleuroploca ) pyrulaeformis Hoernes, Auinger, 1980 [ sic ]— Nikolov 1994: 54 , pl. 5, figs 1–2. Fasc [ iolaria ]. pirulaeformis [ sic ] Hoern. et Auinger— Cossmann 1901: 40 . Type material. Lectotype (designated by Nikolov 1994: 54 ): NHMW 1860 /0040/0177, SL: 105.6 mm , MD : 55.4 mm , Lăpugiu de Sus ( Romania ), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 5), Figs 5A 1 –A 2 . Paralectotypes : NHMW 1866 /0040/0301a, SL: 76.0 mm , MD : 34.3 mm , Lăpugiu de Sus ( Romania ), Figs 5B 1 –B 2 . NHMW 1857 /0024/0021, SL: 95.4 mm , MD : 47.5 mm , Lăpugiu de Sus ( Romania ), Fig. 5C . NHMW 1866 /0040/0301, SL: 55.3 mm , MD : 26.5 mm , Lăpugiu de Sus ( Romania ), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 6) . Additional material. 1 spec. , NHMW 2023 /0367/0001, Baden-Sooss ( Austria ) . Revised description. Very large, broad fusiform shell of six teleoconch whorls with gradate spire; apical angle ~50°. Protoconch unknown. Early teleoconch whorls convex with shallow, concave subsutural ramp. Broad axial ribs forming blunt nodes along periphery, fading on subsutural ramp. Spiral sculpture of several prominent spiral cords, separated by narrower interspaces. Cords prominent on subsutural ramp and between nodes, subobsolete over nodes. Suture moderately incised, with narrow subsutural collar, bearing two spiral cords. Growth lines more prominent on subsutural ramp and strongly raised between spiral cords of subsutural collar. Nodes at periphery split on fourth and fifth teleoconch whorls by intercalation of narrow spiral cord. Last whorl attaining 75% of total height, with broad, shallow, concave subsutural ramp, roundly angled at strong shoulder, convex below, moderately constricted at base; fasciole moderately broad, poorly delimited. Spiral sculpture of about four narrow, widely spaced spirals over subsutural ramp, five spiral rows of large, low nodes below shoulder; interspaces between spiral rows of nodes widening abapically, bearing one or two secondary cords; cords over base prominent, elevated. Aperture moderately wide, pyriform. Columellar callus forming broad thickened rim along siphonal canal, thinner and adherent above. Columella moderately and broadly excavated with three strongly oblique columellar folds deep inside aperture; only two folds visible in aperture of fully-grown specimens; adapical fold weak, abapical fold marking slightly angled transition to siphonal canal. Anal canal narrowly incised. Outer lip thin, distinctly notched at shoulder, with numerous thin lirae extending deep within aperture. Siphonal canal moderately long, narrow, straight or slightly deflected to the left. FIGURE 4. Columellar features of Paratethyan Fasciolariidae . A. Dulaiania pleurotomoides (Hoernes &Auinger 1890) , holotype, NHMW 1855/0043/0041. B. Polygona vindobonensis ( Csepreghy-Meznerics, 1956 ) , NHMW 1860/001/0245, Steinebrunn (Austria). C. Eurolatirus ornatus (d’Orbigny, 1852) , NHMW 2023/0351/0004, Enzesfeld (Austria). D. Tarantinaea hoernesii (Seguenza, 1875) , NHMW 1846/0037/0279, lectotype, Steinebrunn (Austria). E. Neolatirus recticauda (Fuchs in Karrer, 1877), holotype, NHMW 1883/0002/4168, Rauchstallbrunn-Graben at Baden (Austria). F. Cryptoneolatirus bellardii (Hörnes, 1854) , NHMW 1865/0001/0211, Lăpugiu de Sus (Romania). G. Turrilatirus moravicus (Hoernes & Auinger, 1890) , NHMW 1866/0001/0860e, paralectotype, Niederleis (Austria). H. Angustifusus hoessii ( Naumann, 1852 ) , NHMW 2023/0367/0002, Baden-Sooss (Austria). I. Angustifusus vindobonensis (Hoernes & Auinger, 1890) , NHMW 1846/0037/0262a. J. Fraudifusinus crispoides ( Kittl, 1887 ) , NHMW 2023/0350/0009, Bad Vöslau (Austria). K. Fraudifusinus crispoides ( Kittl, 1887 ) , NHMW 2023/0350/0010, Bad Vöslau (Austria). L. Fraudifusinus pseudocrispoides nov. sp. , NHMW 1870/0033/0102, paratype, Lăpugiu de Sus (Romania). M. Fraudifusinus grundensis nov. sp. , NHMW 1855/0045/0720b, holotype, Grund (Austria). N. Pseudolatirus bilineatus ( Naumann, 1852 ) , NHMW 1846/0037/0264e, neotype, Möllersdorf (Austria). O. Takashius boettgeri ( Kovács, 2022 ) , NHMW A1426, Lăpugiu de Sus (Romania). P. Takashius vinculum nov. sp. , NHMW 1868/0001/0083, paratype, Möllersdorf (Austria). FIGURE 5. Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890) . A 1 –A 2 . NHMW 1860/0040/0177, lectotype, Lăpugiu de Sus (Romania). B 1 –B 2 . NHMW 1866/0040/0301a, paralectotype, Lăpugiu de Sus (Romania). C. NHMW 1857/0024/0021, paralectotype, Lăpugiu de Sus (Romania). D 1 –D 2 . Aurantilaria rudis ( Schaffer, 1912 ) , NHMW 1861/0050/0289, lectotype, Gauderndorf (Austria). E 1 –E 2 . Aurantilaria rudis ( Schaffer, 1912 ) , Collection Anton Breitenberger, Mörtersdorf (Austria). Discussion. This species was placed in Aurantilaria by Snyder et al . (2012: 46) , which is followed herein. The early teleoconch whorls of Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890) are nearly indistinguishable from those of Aurantilaria tarbelliana ( Grateloup, 1845 ) , but later whorls differ in their bifid nodes along the periphery. In addition, the spiral rows of nodes on the last whorl distinguish this species from A. tarbelliana , which has a longer siphonal canal. We are not aware of any closely related species in the European Neogene. So far, A. pyrulaeformis is only known from southern basins of the Central Paratethys Sea and was restricted to the early/middle Badenian (Langhian). Aurantilaria carminamaris ( Maury, 1917 ) , from the Messinian Cercado Formation in the Dominican Republic, is a Western Atlantic counterpart of the Paratethyan species but differs in the more pointed nodes and the presence of a single row of nodes along the shoulder (see Landau & Vermeij 2014 : pl. 4, figs 21–25). Paleoenvironment. Unknown; probably middle to outer neritic environments. Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin : Lăpugiu de Sus ( Romania ) (Hoernes & Auinger 1890); Dacian Basin : Bivolare ( Bulgaria ) ( Nikolov 1994 ).