<document id="F34D66B4D9D8C0BB98D5BE06CCC4958A" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. &amp; Kelso, Scott" docDate="2021" docId="03A887D05F61FFAAFF3EEBD5FBEB4E9E" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella helwanensis" docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="36" updateTime="1699012478263" updateUser="plazi">
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<mods:title id="CF5518985869933EA93DE849A452357C">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
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<mods:namePart id="C0209CF5825DADDF940913CCF101E05E">Motamedinia, Behnam</mods:namePart>
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<mods:namePart id="8A4DA351B202953008D8DD75451B73B2">Skevington, Jeffrey H.</mods:namePart>
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<mods:namePart id="83FCC5BE68410B328528E815C558F681">Kelso, Scott</mods:namePart>
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<mods:date id="78D5F54157A74168B5D27BFCFA584F02">2021</mods:date>
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<paragraph id="8BBE36C65F61FFAAFF3EEBD5FD3D4F81" blockId="35.[151,658,151,214]" box="[151,658,151,178]" pageId="35" pageNumber="36">
<heading id="D0F681AA5F61FFAAFF3EEBD5FD3D4F81" bold="true" box="[151,658,151,178]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<taxonomicName id="4C014D455F61FFAAFF3EEBD5FD3D4F81" ID-CoL="7CH4L" authority="(Collin, 1949)" baseAuthorityName="Collin" baseAuthorityYear="1949" box="[151,658,151,178]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="helwanensis">
<emphasis id="B975EAD45F61FFAAFF3EEBD5FD3D4F81" bold="true" box="[151,658,151,178]" pageId="35" pageNumber="36">
<emphasis id="B975EAD45F61FFAAFF3EEBD5FE4E4F82" bold="true" box="[151,481,151,178]" italics="true" pageId="35" pageNumber="36">Tomosvaryella helwanensis</emphasis>
(
<bibRefCitation id="EF904B375F61FFAAFE58EBD5FD254F81" author="Collin, J. E." box="[497,650,151,177]" pageId="35" pageNumber="36" pagination="175 - 225" refId="ref56517" refString="Collin, J. E. (1949) Results of the Armstrong College Expedition to Siwa Oasis (Libyan Desert), 1935, under the leadership of Prof. J. Omer-Cooper. Diptera Empididae, Dolichopodidae, Aschiza and Acalypterae. Bulletin de la Societe Fouad 1 er d'Entomologie, 33, 175 - 225." type="journal article" year="1949">Collin, 1949</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F61FFAAFF3EEBF9FF484FE6" box="[151,231,187,214]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BBE36C65F61FFAAFF3EEBF9FF484FE6" blockId="35.[151,658,151,214]" box="[151,231,187,214]" pageId="35" pageNumber="36">
<figureCitation id="133A2A435F61FFAAFF3EEBF9FF484FE6" box="[151,231,187,214]" captionStart="FIGURE 53" captionStartId="87.[151,250,847,871]" captionTargetBox="[151,1436,181,823]" captionTargetId="figure-21@87.[151,1436,181,823]" captionTargetPageId="87" captionText="FIGURE 53. Distribution of Tomosvaryella ellipiensis sp. nov., T. geniculata, T. helwanensis, T. jubata, T. kiansiae sp. nov., T. nigronitida in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122777" httpUri="https://zenodo.org/record/5122777/files/figure.png" pageId="35" pageNumber="36">Fig. 53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F61FFAAFF3EEA41FECF4EB9" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BBE36C65F61FFAAFF3EEA41FECF4EB9" blockId="35.[151,1437,259,430]" pageId="35" pageNumber="36">
<emphasis id="B975EAD45F61FFAAFF3EEA41FEBB4E2D" bold="true" box="[151,276,259,285]" pageId="35" pageNumber="36">Diagnosis:</emphasis>
Following 
<bibRefCitation id="EF904B375F61FFAAFE3CEA41FD304E2D" author="Kehlmaier, C. &amp; Gibbs, D. J. &amp; Withers, P." box="[405,671,259,285]" pageId="35" pageNumber="36" pagination="31 - 60" refId="ref57273" refString="Kehlmaier, C., Gibbs, D. J. &amp; Withers, P. (2019) New records of big-headed flies (Diptera: Pipunculidae) from the Mediterranean Basin. Bonn Zoological Bulletin, 68 (1), 31 - 60" type="journal article" year="2019">
Kehlmaier 
<emphasis id="B975EAD45F61FFAAFDBBEA46FDE44E2D" box="[530,587,259,285]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
(2019)
</bibRefCitation>
, this species can be recognized by the symmetrical surstyli in dorsal view, broadened at the base, both surstyli slightly bean-shaped in lateral view; epandrium wider than long; phallus trifid, one of ejaculatory duct with small spines; phallic guide bent towards surstyli in lateral view (
<bibRefCitation id="EF904B375F61FFAAFB4FEA09FF7C4EB9" author="Kehlmaier, C. &amp; Gibbs, D. J. &amp; Withers, P." pageId="35" pageNumber="36" pagination="31 - 60" refId="ref57273" refString="Kehlmaier, C., Gibbs, D. J. &amp; Withers, P. (2019) New records of big-headed flies (Diptera: Pipunculidae) from the Mediterranean Basin. Bonn Zoological Bulletin, 68 (1), 31 - 60" type="journal article" year="2019">
Kehlmaier 
<emphasis id="B975EAD45F61FFAAFACCEA0EFA324E55" box="[1381,1437,331,357]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2019
</bibRefCitation>
: fig. 8A–D).
</paragraph>
</subSubSection>
<subSubSection id="C31B654D5F61FFAAFF6EEAD1FBEB4E9E" box="[199,1092,403,430]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BBE36C65F61FFAAFF6EEAD1FBEB4E9E" blockId="35.[151,1437,259,430]" box="[199,1092,403,430]" pageId="35" pageNumber="36">
<emphasis id="B975EAD45F61FFAAFF6EEAD1FECC4E9D" bold="true" box="[199,355,403,429]" pageId="35" pageNumber="36">Distribution:</emphasis>
<collectingCountry id="F31676565F61FFAAFEC3EAD6FE1F4E9E" box="[362,432,404,430]" name="Egypt" pageId="35" pageNumber="36">Egypt</collectingCountry>
, 
<collectingCountry id="F31676565F61FFAAFE12EAD1FE564E9D" box="[443,505,403,429]" name="Israel" pageId="35" pageNumber="36">Israel</collectingCountry>
(Kehlamier 
<emphasis id="B975EAD45F61FFAAFD2FEAD6FD104E9D" box="[646,703,403,429]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2019; 
<bibRefCitation id="EF904B375F61FFAAFCA5EAD1FC624E9D" author="Skevington, J. H." box="[780,973,403,429]" pageId="35" pageNumber="36" refId="ref58350" refString="Skevington, J. H. (2020) &quot; Pipunculidae &quot;. Canadian National Collection of Insects, Arachnids and Nematodes database. Available from: https: // cnc. agr. gc. ca / taxonomy / Taxonomy. php? id = 18557 (accessed 18 October 2020) [public version of Pipunculidae catalogue maintained by J. H. Skevington]" type="url" year="2020">Skevington 2020</bibRefCitation>
) (
<figureCitation id="133A2A435F61FFAAFC4AEAD1FB984E9E" box="[995,1079,403,430]" captionStart="FIGURE 53" captionStartId="87.[151,250,847,871]" captionTargetBox="[151,1436,181,823]" captionTargetId="figure-21@87.[151,1436,181,823]" captionTargetPageId="87" captionText="FIGURE 53. Distribution of Tomosvaryella ellipiensis sp. nov., T. geniculata, T. helwanensis, T. jubata, T. kiansiae sp. nov., T. nigronitida in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122777" httpUri="https://zenodo.org/record/5122777/files/figure.png" pageId="35" pageNumber="36">Fig. 53</figureCitation>
).
</paragraph>
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</treatment>
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