New data on the Harpacteinae of Turkey (Araneae, Dysderidae) Author Kunt, Kadir Boğaç Department of Biology, Faculty of Science, Eskişehir Technical University, TR- 26470 Eskişehir, Turkey & Cyprus Wildlife Research Institute, Taşkent, Kyrenia, Cyprus Author Özkütük, Recep Sulhi Department of Biology, Faculty of Science, Eskişehir Technical University, TR- 26470 Eskişehir, Turkey text Zootaxa 2023 2023-11-23 5375 3 379 408 https://mapress.com/zt/article/download/zootaxa.5375.3.4/52335 journal article 10.11646/zootaxa.5375.3.4 1175-5326 10199980 26591537-2135-4301-AD46-FF4D2D9F7C68 Harpactea elvericii sp. nov. Figs 37–43 Material examined: Holotype 1 ♂ ( ETZM ), Alanya Dist. , Avsallar Town ( 36°38’21.50”N 31°45’24.88”E ), asl c. 30 m , 04 Jan 2013 , leg. M. Elverici. Paratypes 5 ♂ , 4 ♀ , 1J ( ETZM ), same data as holotype — 1 ♂ , 2 ♀ ( ETZM ), Alanya Dist. , Avsallar Town ( 36°38’21.50”N 31°45’24.88”E ), asl c. 30 m , 04 Jan 2013 , leg. Kadir Boğac Kunt — 1 ♀ , 1 ♂ , 2J ( ETZM ), Alanya Dist. , Dim Dam ( 36°33’48.48”N 32°9’4.42”E ), asl c. 226 m , 06 Jan 2013 , leg. Kadir Boğac Kunt — 3 ♂ , 4 ♀ ( ETZM ), Alanya Dist. , Dim Valley ( 36°32’28.00”N 32° 5’49.00”E ), asl c. 44 m , 6 Jan 2013 , leg. M. Elverici — 2 ♀ , 8J ( ETZM ), Alanya Dist. , Dim Valley ( 36°32’33.15”N 32° 6’18.28”E ), asl c. 20 m , 15 Sept 2013 , leg. Kadir Boğac Kunt — 4 ♂ , 6 ♀ ( ETZM ), GazipaŞa Dist. , Adanda Vill. , Adanda Castle ( 36°14’13.00”N 32°28’4.00”E ), asl c. 800 m , 29 Mar 2017 , leg. Kadir Boğac Kunt. Etymology: The specific epithet is a patronym, honoring a friend of the authors, Turkish arachnologist, Dr. Mert Elverici. Diagnosis: The male of H. elvericii sp. nov. closely resembles H. bilecenoglui sp. nov. by the lack of conductor and the straigh embolus, but they can be easily distinguished by the shape of the embolus ( Figs 27–30 , 40–41 ). Namely, in H. bilecenoglui sp. nov. the embolus becomes distally thinner, after the proximal half, whereas in H. elvericii sp. nov. becomes thinner after the first proximal quarter. The bulb of both species is also very similar to that of H. sanctaeinsulae ( Figs 44–45 ), however, in the latter, the embolus is thinner and more delicate. The vulva of H. elvericii sp. nov. resembles that of H. prope sbordonii Brignoli, 1978 by the short and folded spermatheca, but it differs by the very short and spine-like spermathecal keel in H. elvericii sp. nov. , while in H. prope sbordonii is longer and square-like outgrowth at the frontal margin ( Fig. 43 ). Description: Measurements: [ Holotype ♂ / Paratypes ♀ ] TL 4.35/4.90–5.20; AL 2.35/2.75–3.00; CL 2.00/2.15–2.20; CW 1.55/1.60–1.65; Clh 0.06/0.04–0.06; AEd 0.12/0.12–0.12; iAE 0.04/0.03–0.04; PLEd 0.11/0.11–0.11; PMEd 0.09/0.10–0.10; ChF 0.23/0.23–0.35; ChG 0.23/0.23–0.26; ChL 0.70/0.75–0.80. [ Paratype ♂ ] TL 5.05; AL 2.90; CL 2.15; CW 1.65; Clh 0.05; AEd 0.12; iAE 0.02; PLEd 0.11; PMEd 0.10; ChF 0.25; ChG 0.25; ChL 0.85. Medium sized harpacteine spiders. Carapace reddish in both sexes. There is no difference in tone between the head and thoracic region. Thoracic regions pentagonal; head region short, narrow and rounded anteriorly. Fovea dark brown, longitudinal and distinct ( Fig. 37 ). AE, PLE and PME close to each other, arranged annularly, silvery colored. Sternum, labium, gnahtocoxae brown. Sternum yellowish with brownish setae. Tips of the labium and gnatocoxae with blackish setae. Chelicerae reddish. Surface of the chelicerae medially covered with protuberances bearing short black setae. Cheliceral groove with four weakly developed teeth. Promarginal teeth are larger than retromarginal teeth. Of the promarginal teeth, the first tooth near the base of the cheliceral groove is almost half as large as the second. There is a gap between the two about the width of the base of the second tooth. The first retromarginal tooth is in the centre of the cheliceral groove and aligned between the promarginal teeth. The second retromarginal tooth on top is almost twice the size of the first ( Fig. 39 ). Abdomen grayish, cylindrical. Surface covered with blackish setae. Some males with a scutum indistinctly, others with a very distinct scutum ventrally on the abdomen, pregastrally positioned on the right and left ( Fig. 38 ). Legs yellowish brown. Anterior legs darker than posteriors. Leg formula: VI, I, II, III. Leg spination and measurements given in Tables 9 and 10. FIGURES 37-39. Harpactea elvericii sp. nov. 37. Male, habitus 38. Ditto, anterior ventral area 39. Chelicerae, ventral view (red dots, promarginal; white dots, retromarginal teeth). Abbrevation: ES , epigastral scutum. FIGURES 40–43. Harpactea elvericii sp. nov. 40. Bulb (left), prolateral view 41. Bulb (right), retrolateral view 42. Vulva, dorsal view 43. Ditto, nearly dorsal view. Abbrevations: AA , anterior margin of the anterior arch; AC , anterior arc; E , embolus; PD , posterior diverticulum; S , spermatheca; SK , spermathecal keel; T , tegulum; TB , transversal bar.

Legs	I	II	III	IV
♂
Fe	1.75–2.00	1.60–1.75	1.30–1.40	1.85–2.00
Pa	1.00–1.25	1.00–1.15	0.60–0.75	1.00–1.00
Ti	1.40–1.50	1.25–1.50	0.90–1.00	1.50–1.60
Me	1.25–1.50	1.10–1.50	1.00–1.25	1.50–1.65
Ta	0.40–0.50	0.30–0.45	0.30–0.30	0.40–0.50
Total	5.80–6.75	5.25–6.35	4.10–4.70	6.25–6.75
♀
Fe	1.65–1.75	1.50–1.50	1.00–1.30	1.80–2.00
Pa	1.15–1.15	1.00–1.00	0.50–0.70	0.90–1.00
Ti	1.35–1.40	1.00–1.00	1.00–1.00	1.50–1.65
Me	1.25–1.25	1.15–1.25	1.05–1.25	1.75–1.75
Ta	0.35–0.40	0.35–0.40	0.40–0.40	0.45–0.50
Total	5.75–5.95	5.00–5.15	3.95–4.65	6.40–6.90

FIGURES 44–45. Harpactea sanctaeinsulae , holotype 39. Bulb, prolateral view 40. Ditto, retrolateral view. Palp ( Figs 40–41 ): Palpal tarsus longer than tibia. Tegulum oval, more rounded posteriorly than anteriorly. Embolus spiniform, broad at base, roughly the same length as tegulum. Embolus displays a very slight dorsal sinuosity medially. Conductor and median apophysis absent. Vulva ( Figs 42–43 ): Spermatecha characteristic; extremely broad anteriorly.We can suggest that the spermatheca structurally consists of three parts. The distal, widest first part, to which the spermathecal keel is attached ( Fig. 43 ). Below it, in the centre, is the second part, almost half the width of the first. Between the first and second parts, there is a deep cleft ( Fig. 42 ). The third part merges with the roundish structures and is rod-shaped. Spermathecal keel reduced to a very short spike on the left side. Transversal bar in the form of a thin belt, the middle where it joins the receptaculum is relatively well sclerotised. Posterior diverticulum rounded, distinct ( Fig. 42 ).