<document id="CC0FD3D6AA03BF89A599D720D32A687F" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFDEBB09FF188709FF61FB16" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia troglodytes Gompel 2020, comb. nov." docType="treatment" docVersion="7" lastPageNumber="175" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="175" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4E6755BD6356E1536CB7B3437F798D99">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="C3B5FAEFB84D276EFE9D394920CD6ADA">Gompel, Nicolas</mods:namePart>
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<mods:date id="EB5755A82E8A199082D5EA7D1DEA5FBE">2020</mods:date>
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<treatment id="039187D8FFDEBB09FF188709FF61FB16" ID-DOI="http://doi.org/10.5281/zenodo.4453466" ID-GBIF-Taxon="177012495" ID-Zenodo-Dep="4453466" LSID="urn:lsid:plazi:treatment:039187D8FFDEBB09FF188709FF61FB16" httpUri="http://treatment.plazi.org/id/039187D8FFDEBB09FF188709FF61FB16" lastPageNumber="175" pageId="24" pageNumber="175">
<subSubSection id="C3226545FFDEBB09FF188709FC97FCA2" box="[151,778,834,861]" pageId="24" pageNumber="175" type="nomenclature">
<paragraph id="8B8736CEFFDEBB09FF188709FC97FCA2" blockId="24.[151,778,834,897]" box="[151,778,834,861]" pageId="24" pageNumber="175">
<heading id="D0CF81A2FFDEBB09FF188709FC97FCA2" bold="true" box="[151,778,834,861]" fontSize="11" level="1" pageId="24" pageNumber="175" reason="1">
<emphasis id="B94CEADCFFDEBB09FF188709FC97FCA2" bold="true" box="[151,778,834,861]" pageId="24" pageNumber="175">
<taxonomicName id="4C384D4DFFDEBB09FF188709FDEBFCA3" authority="(Champion, 1916)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Champion" baseAuthorityYear="1916" box="[151,630,834,861]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="troglodytes" status="comb. nov.">
<emphasis id="B94CEADCFFDEBB09FF188709FE11FCA2" bold="true" box="[151,396,834,861]" italics="true" pageId="24" pageNumber="175">Zarcosia troglodytes</emphasis>
(Champion, 1916)
</taxonomicName>
, 
<taxonomicNameLabel id="A27F57A7FFDEBB09FD0C8709FC97FCA2" box="[643,778,835,861]" pageId="24" pageNumber="175" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF18872DFEE1FC7E" box="[151,380,871,897]" pageId="24" pageNumber="175" type="description">
<paragraph id="8B8736CEFFDEBB09FF18872DFEE1FC7E" blockId="24.[151,778,834,897]" box="[151,380,871,897]" pageId="24" pageNumber="175">
(
<figureCitation id="13032A4BFFDEBB09FF10872DFE96FC7E" box="[159,267,871,897]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="24" pageNumber="175">Figures 1</figureCitation>
; 
<figureCitation id="13032A4BFFDEBB09FE97872DFED6FC7E" box="[280,331,871,897]" captionStart="FIGURE 2" captionStartId="8.[151,250,1960,1984]" captionTargetBox="[170,1419,212,1927]" captionTargetId="figure-30@8.[153,1434,178,1933]" captionTargetPageId="8" captionText="FIGURE 2. continued next page." figureDoi="http://doi.org/10.5281/zenodo.4403789" httpUri="https://zenodo.org/record/4403789/files/figure.png" pageId="24" pageNumber="175">2a,b</figureCitation>
; 
<figureCitation id="13032A4BFFDEBB09FED8872DFEE9FC7E" box="[343,372,871,897]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="24" pageNumber="175">4q</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF1887FAFD5CFC13" pageId="24" pageNumber="175" type="reference_group">
<paragraph id="8B8736CEFFDEBB09FF1887FAFC8BFC37" blockId="24.[151,1018,944,1040]" box="[151,790,944,969]" pageId="24" pageNumber="175">
<taxonomicName id="4C384D4DFFDEBB09FF1887FAFDC9FC36" authority="Champion, 1916: 30" authorityName="Champion" authorityPageNumber="30" authorityYear="1916" box="[151,596,944,969]" class="Insecta" family="Aderidae" genus="Xylophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="troglodytes">
<emphasis id="B94CEADCFFDEBB09FF1887FAFEE6FC37" box="[151,379,944,968]" italics="true" pageId="24" pageNumber="175">Xylophilus troglodytes</emphasis>
Champion, 1916: 30
</taxonomicName>
; 
<typeStatus id="5483886CFFDEBB09FDD187FAFD5DFC37" box="[606,704,944,968]" pageId="24" pageNumber="175" type="holotype">Holotype</typeStatus>
: BMNH
</paragraph>
<paragraph id="8B8736CEFFDEBB09FF18879EFD5CFC13" blockId="24.[151,1018,944,1040]" box="[151,705,980,1004]" pageId="24" pageNumber="175">
<taxonomicName id="4C384D4DFFDEBB09FF18879EFDADFC13" authority="(Champion, 1916)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Champion" baseAuthorityYear="1916" box="[151,560,980,1004]" class="Insecta" family="Aderidae" genus="Euglenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="troglodytes" status="comb. nov.">
<emphasis id="B94CEADCFFDEBB09FF18879EFEF6FC13" box="[151,363,980,1004]" italics="true" pageId="24" pageNumber="175">Euglenes troglodytes</emphasis>
(Champion, 1916)
</taxonomicName>
: 
<bibRefCitation id="EFA94B3FFFDEBB09FDB7879FFD5CFC13" author="Werner, F. G." box="[568,705,980,1004]" pageId="24" pageNumber="175" pagination="121 - 127" refId="ref18162" refString="Werner, F. G. (1962) The Aderidae of the Batu Caves, Malaya (Coleoptera). Pacific Insects, 4, 121 - 127." type="journal article" year="1962">Werner, 1962</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF1887B3FC67FBEF" box="[151,1018,1016,1040]" pageId="24" pageNumber="175" type="description">
<paragraph id="8B8736CEFFDEBB09FF1887B3FC67FBEF" blockId="24.[151,1018,944,1040]" box="[151,1018,1016,1040]" pageId="24" pageNumber="175">
<taxonomicName id="4C384D4DFFDEBB09FF1887B3FDB1FBEF" authority="(Champion, 1916)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Champion" baseAuthorityYear="1916" box="[151,556,1016,1040]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="troglodytes">
<emphasis id="B94CEADCFFDEBB09FF1887B3FEF5FBEF" box="[151,360,1016,1040]" italics="true" pageId="24" pageNumber="175">Zarcosia troglodytes</emphasis>
(Champion, 1916)
</taxonomicName>
: unpublished combination in 
<bibRefCitation id="EFA94B3FFFDEBB09FCD987B2FC67FBEF" author="Grzymala, T. L." box="[854,1018,1016,1040]" pageId="24" pageNumber="175" refId="ref17990" refString="Grzymala, T. L. (2016) Systematics of Puppet Beetles (Coleoptera, Aderidae): Classification, Molecular Phylogenetics &amp; Evolution of Sexual Dimorphism. Environmental Science, Policy, and Management. University of California, Berkeley, California, 409 pp." type="book" year="2016">Grzymala, 2016</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF188075FB9CFBA6" box="[151,1025,1086,1113]" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="8B8736CEFFDEBB09FF188075FB9CFBA6" blockId="24.[151,1436,1086,1257]" box="[151,1025,1086,1113]" pageId="24" pageNumber="175">
<materialsCitation id="3B503C93FFDEBB09FF188075FC60FBA6" ID-GBIF-Occurrence="3020160420" box="[151,1021,1086,1113]" country="Malaysia" county="Darul Ehsan" location="Batu" municipality="Gombak District" pageId="24" pageNumber="175" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="B94CEADCFFDEBB09FF188075FEABFBA7" bold="true" box="[151,310,1086,1113]" pageId="24" pageNumber="175">
<typeStatus id="5483886CFFDEBB09FF188075FF4FFBA6" box="[151,210,1087,1113]" pageId="24" pageNumber="175">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFDEBB09FEB38075FE3AFBA6" box="[316,423,1087,1113]" name="Malaysia" pageId="24" pageNumber="175">Malaysia</collectingCountry>
: 
<collectingRegion id="49FCF82CFFDEBB09FE3D8075FD8BFBA6" box="[434,534,1087,1113]" country="Malaysia" name="Selangor" pageId="24" pageNumber="175">Selangor</collectingRegion>
<collectingCounty id="62E64E42FFDEBB09FD938075FD37FBA6" box="[540,682,1087,1113]" pageId="24" pageNumber="175">Darul Ehsan</collectingCounty>
, 
<collectingMunicipality id="6BE3ACB4FFDEBB09FD3A8075FCE9FBA6" box="[693,884,1087,1113]" pageId="24" pageNumber="175">Gombak District</collectingMunicipality>
, 
<location id="8EE76015FFDEBB09FCF08075FC29FBA6" LSID="urn:lsid:plazi:treatment:039187D8FFDEBB09FF188709FF61FB16:8EE76015FFDEBB09FCF08075FC29FBA6" box="[895,948,1087,1113]" country="Malaysia" county="Darul Ehsan" municipality="Gombak District" name="Batu" pageId="24" pageNumber="175" stateProvince="Selangor">Batu</location>
caves
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF488028FD5FFB82" box="[199,706,1122,1149]" pageId="24" pageNumber="175" type="distribution">
<paragraph id="8B8736CEFFDEBB09FF488028FD5FFB82" blockId="24.[151,1436,1086,1257]" box="[199,706,1122,1149]" pageId="24" pageNumber="175">
<emphasis id="B94CEADCFFDEBB09FF488028FEFCFB83" bold="true" box="[199,353,1122,1148]" pageId="24" pageNumber="175">Distribution.</emphasis>
Malaya, Batu caves (
<figureCitation id="13032A4BFFDEBB09FDDD8029FD2CFB82" box="[594,689,1123,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="24" pageNumber="175">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF4880CCFF61FB16" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="8B8736CEFFDEBB09FF4880CCFF61FB16" blockId="24.[151,1436,1086,1257]" pageId="24" pageNumber="175">
<emphasis id="B94CEADCFFDEBB09FF4880CCFE2FFB5F" bold="true" box="[199,434,1158,1184]" pageId="24" pageNumber="175">Material examined.</emphasis>
<materialsCitation id="3B503C93FFDEBB09FE3580CCFE2DFB3A" ID-GBIF-Occurrence="3020160429" collectionCode="BMNH" collectorName="H. N. Ridley" country="Malaysia" location="Gombak District" pageId="24" pageNumber="175" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Selangor">
<emphasis id="B94CEADCFFDEBB09FE3580CCFDBAFB5F" bold="true" box="[442,551,1158,1184]" pageId="24" pageNumber="175">
<collectingCountry id="F32F765EFFDEBB09FE3580CCFDBAFB5F" box="[442,551,1158,1184]" name="Malaysia" pageId="24" pageNumber="175">Malaysia</collectingCountry>
</emphasis>
: 
<collectingRegion id="49FCF82CFFDEBB09FDB880CDFD06FB5E" box="[567,667,1159,1185]" country="Malaysia" name="Selangor" pageId="24" pageNumber="175">Selangor</collectingRegion>
Darul Ehsan, 
<location id="8EE76015FFDEBB09FCB180CDFC63FB5E" LSID="urn:lsid:plazi:treatment:039187D8FFDEBB09FF188709FF61FB16:8EE76015FFDEBB09FCB180CDFC63FB5E" box="[830,1022,1159,1185]" country="Malaysia" name="Gombak District" pageId="24" pageNumber="175" stateProvince="Selangor">Gombak District</location>
, Batu caves (
<collectorName id="26CD5318FFDEBB09FB1880CDFAB7FB5D" box="[1175,1322,1159,1186]" pageId="24" pageNumber="175">H. N. Ridley</collectorName>
) (
<specimenCount id="9D3EFD47FFDEBB09FACA80CDFAF2FB5E" box="[1349,1391,1159,1185]" pageId="24" pageNumber="175" type="male">1 ♂</specimenCount>
holotype and 
<specimenCount id="9D3EFD47FFDEBB09FE9A80E1FEDDFB3A" box="[277,320,1195,1221]" pageId="24" pageNumber="175" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED29AE0BFFDEBB09FEC080E1FE35FB3A" box="[335,424,1195,1221]" pageId="24" pageNumber="175">BMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B503C93FFDEBB09FE3380E1FF65FB16" ID-GBIF-Occurrence="3020160388" collectingDate="1926-07" collectionCode="BMNH" country="Malaysia" location="Gombak District" pageId="24" pageNumber="175" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Selangor">
<collectingRegion id="49FCF82CFFDEBB09FE3380E1FDBDFB3A" box="[444,544,1195,1221]" country="Malaysia" name="Selangor" pageId="24" pageNumber="175">Selangor</collectingRegion>
<collectorName id="26CD5318FFDEBB09FDA880E1FD28FB3A" box="[551,693,1195,1221]" pageId="24" pageNumber="175">Darul Ehsan</collectorName>
, 
<location id="8EE76015FFDEBB09FD4D80E1FC1CFB3A" LSID="urn:lsid:plazi:treatment:039187D8FFDEBB09FF188709FF61FB16:8EE76015FFDEBB09FD4D80E1FC1CFB3A" box="[706,897,1195,1221]" country="Malaysia" name="Gombak District" pageId="24" pageNumber="175" stateProvince="Selangor">Gombak District</location>
, 
<collectorName id="26CD5318FFDEBB09FC0280E1FC5FFB3A" box="[909,962,1195,1221]" pageId="24" pageNumber="175">Batu</collectorName>
caves (
<collectorName id="26CD5318FFDEBB09FB9680E1FB1FFB3A" box="[1049,1154,1195,1221]" pageId="24" pageNumber="175">C. Dover</collectorName>
) 
<date id="FF86100EFFDEBB09FB1E80E1FB6BFB3A" box="[1169,1270,1195,1221]" pageId="24" pageNumber="175" value="1926-07">
<collectingDate id="EFC2E9E6FFDEBB09FB1E80E1FB6BFB3A" box="[1169,1270,1195,1221]" pageId="24" pageNumber="175" value="1926-07">VII.1926</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFDEBB09FA8880E1FAAFFB3A" box="[1287,1330,1195,1221]" pageId="24" pageNumber="175" type="male">2 ♂</specimenCount>
and 
<specimenCount id="9D3EFD47FFDEBB09FAE580E1FA08FB3A" box="[1386,1429,1195,1221]" pageId="24" pageNumber="175" type="female">1 ♀</specimenCount>
, 
<collectionCode id="ED29AE0BFFDEBB09FF188085FF6DFB16" box="[151,240,1231,1257]" pageId="24" pageNumber="175">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>