Notes on Benstonea (Pandanaceae) from the islands of Halmahera, New Guinea and Sulawesi Author Callmander, Martin W. Missouri Botanical Garden, P. O. Box 299, St. Louis, MO 63166 - 0299, U. S. A. & Conservatoire et Jardin botaniques de la ville de Genève, ch. de l’Impératrice 1, case postale 60, 1292 Chambésy, Switzerland Author Buerki, Sven Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, U. K. Author Keim, Ary P. Herbarium Bogoriense, Botany Division, Research Center for Biology, Indonesian Institute of Sciences, Cibinong Science Center (CSC), Jl. Raya Jakarta-Bogor Km 46, Cibinong, Bogor 16911, Indonesia Author Phillipson, Peter B. Missouri Botanical Garden, P. O. Box 299, St. Louis, MO 63166 - 0299, U. S. A. & Muséum national d’Histoire Naturelle, Département Systématique et Évolution (UMR 7205), text Phytotaxa 2014 2014-08-08 175 3 161 165 http://dx.doi.org/10.11646/phytotaxa.175.3.6 journal article 10.11646/phytotaxa.175.3.6 1179-3163 5143877 Benstonea verruculosa (Backer ex Stone 1978: 55 ) Callm., Buerki & Phillipson , comb. nov. Basionym : Pandanus verruculosus Backer ex Stone (1978: 55) . Type :— INDONESIA . North Maluku Province : Halmahera , Galileo , Sao Tobaroe , Beguin 1920 ( holotype , BO [2 sheets]!, isotype, L [ L0050725 ] image seen) . Notes: — Benstonea verruculosa was described based on a fragmentary collection from Halmahera dating back to 1921 ( Beguin 1920 ). Its overall morphology bears a strong resemblance to B. stenocarpa but its recent rediscovery on the island and collection of more adequate material ( Callmander & Lasut 1151 ) provides a clarification of its delimitation and ample justification for its recognition at species level. Benstonea verruculosa is characterized by larger and broader leaves (c. 450 x 14 cm ) (400 x 11 cm in B. stenocarapa ), but more significant is the distinct morphology of its drupes. The style of B. verruculosa are short ( Fig. 1A–G ), and not elongated and gradually tapering as in B. stenocarpa . The stigmatic groove is also accordingly much shorter in B. verruculosa . Furthermore, the pileus of B. verruculosa is conspicuously verrucate contrasting with the rather smooth pileus (rarely roughly granular) of B. stenocarpa . The type of B. verruculosa is fragmentary, consisting of only leaf portions and separate drupes. Callmander & Lasut 1151 is the first complete collection of B. verruculosa , a species endemic to Halmahera where it grows along streams. Additional specimens seen :— INDONESIA . North Maluku Province : Halmahera , Lolea Poeloe Rao , 40 m , 12 October 1921 , Beguin 1816 ( BO , L ) ; Soa Tobaroe , 60 m , 2 June 1922 , Beguin 1933 ( BO ) ; Soa Tobaroe , 60 m , 13 August 1922 , Beguin 2119 ( BO ) ; Soa Tobaroe , 60 m , 13 August 1922 , Beguin 2120 ( BO , L ) ; Central Halmahera , Doromesmesan , Loy Poloy , 00°30’29”N , 127°54’21”E , 15 m , 9 February 2013 , Callmander & Lasut 1151 ( BO , G , L , MO ) .