The pholcid spiders of Sri Lanka (Araneae: Pholcidae)
Author
Huber, Bernhard A.
text
Zootaxa
2019
2019-01-23
4550
1
1
57
journal article
27502
10.11646/zootaxa.4550.1.1
291701d4-652e-440e-adab-f6b0201c0972
1175-5326
2625030
2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833
Belisana badulla
sp. n.
Figures 17–22
,
25–26
Belisana
Benj
44:
Eberle
et al.
2018
(molecular data).
Huber
et al.
2018
: fig. 8.
Diagnosis
. Easily distinguished from most congeners by curvature of procursus (towards ventral;
Figs 17, 19
); from two Sri Lankan species with similar procursus (
B. keyti
Huber, 2005
,
B. benjamini
Huber, 2005
) by bulbal apophysis (much larger and wider in
B. keyti
; absent in
B. benjamini
; compare
Figs 22–24
). Females are difficult to distinguish externally from similar congeners; pockets apparently consistently closer together than in
B. keyti
(200– 250 µm versus 260–270 µm; compare
Figs 25–28
); internal genitalia with distinctive pore plates (lateral round part with long narrow elongation towards median;
Fig. 26
; similar only in
B. keyti
); without pair of internal folds as in
B. keyti
(cf.
Fig. 28
).
Etymology
. The specific name is derived from the
type
locality (noun in apposition).
Type material
.
SRI LANKA
:
♂
holotype
,
ZFMK
(
Ar
20001),
Central Province
,
Badulla District
,
Ohiya
(
6.842°N
,
80.885°E
),
1280 m
a.s.l.
,
16.x.2011
(
S.P. Benjamin
)
.
Other material examined
.
SRI LANKA
:
2♂
2♀
,
ZFMK
(
Ar
20002), and
1♂
2♀
6 juvs in pure ethanol,
ZFMK
(
Benj
44), same data as holotype
.
3♀
in pure ethanol,
ZFMK
(
Benj
46), same locality
,
26.v.2012
(S.P. Benjamin, N. Athukorala).
2♂
,
ZFMK
(
Ar
5199, 5396),
Horton Plains
[
6.84°N
,
80.81°E
], 7/
9.iii.2000
(
S.P. Benjamin
)
.
1♂
2♀
,
ZFMK
(
Ar
20003),
Central Province
,
Nuwara Eliya District
,
Horton Plains
N.P. [
6.84°N
,
80.81°E
], ca.
2000 m
a.s.l.
,
20–21.ii.2007
(
S. Benjamin
,
Z. Jaleel
)
.
2♂
4♀
,
ZFMK
(
Ar
5201),
Central Province
,
Nuwara Eliya District
,
Peak Wilderness Sanctuary
[
6.82°N
,
80.50°E
],
22.ii.2007
(
S. Benjamin
,
Z. Jaleel
)
.
2♂
6♀
,
ZFMK
(
Ar
5202),
Agrabopath Forest Reserve
,
Agrapathana
[=Agrapatana-Bopathalawa Forest Reserve,
6.843°N
,
80.678°E
],
vi.2003
(
S.P. Benjamin
)
.
Description. Male
(
holotype
). MEASUREMENTS. Total body length 1.9, carapace width 0.7. Distance PME- PME 140 µm, diameter PME 60 µm, distance PME-ALE 15 µm; AME absent. Sternum width/length: 0.48/0.44. Leg 1: 16.9 (4.2 + 0.3 + 4.1 + 6.7 + 1.6), tibia 2: 2.5, tibia 3: 1.7, tibia 4: 2.4; tibia 1 L/d: 63.
COLOR (in ethanol). Entire spider whitish to pale ochre-yellow, abdomen pale gray, legs without dark rings.
BODY. Habitus as in
B. keyti
(cf.
Fig. 4
); ocular area not raised; carapace without median furrow; clypeus and sternum unmodified.
CHELICERAE. As in
Figs 20–21
, with pair of small apophyses proximally laterally and pair of larger apophyses with hooked tips distally; distance between tips of distal apophyses 0.28.
PALPS. Proximal segments very similar to
B. keyti
(cf. figs
617–618 in
Huber 2005); coxa unmodified; trochanter with short apophysis with ventral and retrolateral rounded processes; procursus as in
Figs 17–19
, membranous distal elements poorly visible in dissecting microscope; bulb as in
Fig. 22
, with distinctive apophysis with curved tip and embolus with subdistal spine-like process.
LEGS. Without spines and curved hairs; few vertical hairs (proximally on metatarsi 1 and
2 in
higher than usual density); retrolateral trichobothrium on tibia 1 at 12%; prolateral trichobothrium apparently absent on tibia 1; tarsus 1 with ~25 pseudosegments, poorly visible in dissecting microscope.
Male
(variation). Tibia
1 in
6 other males: 4.3–4.8 (mean 4.5).
Female
. In general similar to male; tibia
1 in
11 females
: 2.9–3.4 (mean 3.1). Epigynum externally very simple, barely distinguishable from surrounding cuticle, with pair of pockets ~200–250 µm apart, pockets in some females not visible in dissecting microscope. Internal genitalia as in
Figs 25–26
, with distinctive pore plates consisting of round lateral part and long narrow elongation towards median;
Fig. 26
).
Distribution
. Known from several high elevation localities in central
Sri Lanka
(
Fig. 220
).