One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations Author Zamani, Alireza A21C0B82-E2D9-4402-9408-D6160843DAF4 Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa. zamani.alireza5@gmail.com Author Chatzaki, Maria 2FBB48ED-84FE-4386-8F47-C958E9F8A2A1 maria.chatzaki@gmail.com Author Esyunin, Sergei L. 2B591857-5FAB-4B7D-B589-6C4977A3A493 Sergei. Sergei.Esyunin@psu.ru Author Marusik, Yuri M. F215BA2C-5072-4CBF-BA1A-5CCBE1626B08 yurmar@mail.ru text European Journal of Taxonomy 2021 2021-06-04 751 1 68 114 http://dx.doi.org/10.5852/ejt.2021.751.1381 journal article 6279 10.5852/ejt.2021.751.1381 4e3af683-d8a6-40de-991b-c4584d66b8cc 2118-9773 4906007 1D27EA41-2396-463A-B1B2-0CCF79C9F3D2 Marjanus isfahanicus sp. nov. urn:lsid:zoobank.org:act: 9CB7FC88-C04C-4670-A98D-7A3ACC436976 Fig. 16 Diagnosis The new species differs from the generotype by having interrupted anterior margins (vs uninterrupted), a strongly rebordered W-shaped atrium (vs inconspicuous) and round receptacles (vs tubular). Etymology The specific epithet is an adjective and refers to Isfahan Province , Iran , the type locality of the new species. Type material Holotype IRAN • ♀ ; Isfahan Province , Qamsar & Barzok Protected Area , 55 km SW of Qamsar , 14 km NE of Kamoo , near the road of Gargash observatory ; 33°37′ N , 51°19′ E ; 2710 m a.s.l. ; 19 May 2016 ; P. Ponel leg.; MHNG . Description Female Habitus as in Fig. 16A . Total length 3.04. Carapace 1.40 long, 1.01 wide. Eye sizes and interdistance of PMEs: AME = 0.05, ALE = 0.07, PME = 0.06, PLE = 0.06, PME–PME = 0.05. Carapace and chelicerae dark brown; carapace darker at the margins, with a pattern of light patches. Chelicera with 3 promarginal teeth and 1 retromarginal tooth. Sternum, maxillae and labium light brown. Legs lighter than carapace, without annulations, tarsi lighter. Abdomen dark dorsally, light brown ventrally, without any markings. Spinnerets light gray and uniform in color. Measurements of legs: I: 3.27 (0.95, 0.63, 0.69, 0.54, 0.46), II: 3.14 (0.91, 0.52, 0.69, 0.52, 0.50), III: 2.59 (0.74, 0.37, 0.51, 0.47, 0.50), IV: 3.98 (1.08, 0.56, 0.83, 0.92, 0.59). Epigyne as in Fig. 16B–G . Atrium with distinct and separate anterior hoods; lateral margins distinct, converging; endogyne with membranous bursae near copulatory openings; copulatory ducts thin, forming 2 screw-like, full lateral coils in mid part, straight posteriorly, with blind glandular heads, connected Fig. 16. Marjanus isfahanicus sp. nov. , holotype, ♀ (MHNG). A . Habitus, dorsal view. B, D, F . Epigyne, ventral view. C, E, G . Epigyne, dorsal view. Scale bars: A = 0.5 mm; B = 0.2 mm; F–G = 0.1 mm. to ventral additional bursae with extra glandular tubes; receptacles globular, 3 times as wide as ducts, touching each other, with fertilization ducts located at their upper part. Male Unknown. Comments Based on the similarity of the epigynal conformation to that of the type species, we tentatively place this species here in the absence of male specimens. However, only two of the three diagnostic elements of the females of this genus are met in this new species: the long, coiled copulatory ducts and additional anterior copulatory bursae and the receptacles being round rather than tubular. The latter character may have to be omitted from the generic diagnosis when more species are described and included in this genus. Distribution Known only from the type locality in Isfahan Province , central Iran . This is the first record of Marjanus in Iran , and the second species described in this genus.