The first fossil buprestids from the Middle Jurassic Jiulongshan Formation of China (Coleoptera: Buprestidae)
Author
Pan, Xiaoxiong
Author
Chang, Huali
Author
Ren, Dong
Author
Shih, Chungkun
text
Zootaxa
2011
2745
53
62
journal article
10.5281/zenodo.276648
21fd10d3-4445-4780-912e-282cddd0f36c
1175-5326
276648
Sinoparathyrea robusta
Pan, Chang & Ren
,
sp. nov.
(
Figs. 8, 9
,
12, 13
)
Type
specimen.
Holotype
, male, CNU-COL-NN2010410 PC (part and counterpart of one fossil specimen), housed in the Key Lab of Insect Evolution & Environmental Changes, Capital Normal University, Beijing,
China
.
Locality and horizon.
Daohugou Village, Middle Jurassic Jiulongshan Formation, Ningcheng County, Inner
Mongolia
,
China
.
Etymology.
The species name is derived from the Latin word “
robustus
”, referring to its wider body.
Diagnosis.
Body wide; pronotum wider, about 1.65 times as wide as long, anterior pronotal margin straight, posterior margin slightly bisinuate; Elytra 3.0 times as long as wide, disc with nine distinct longitudinal striae, the 6th and 7th striae fading out at 1/5 of elytron length from apex; the first ventrite 2.5 times as long as the second one.
Description.
Body medium sized,
13.6 mm
long and
6.7 mm
wide, subcylindrical; pronotum slightly narrower than elytra; surface of body with uniform, small, rounded and dense punctures (
Figs. 8
,
12
).
Head: frontovertex convex, surface with granulose and coarse punctate dorsally; eyes relatively small, oval (
Fig. 8
).
FIGURES 8−9.
Sinoparathyrea robusta
sp. nov.
,
line drawings of the holotype. 8—dorsal view, No. CNU-COL- NN2010410P; 9—ventral view, CNU-COL-NN2010410C.
Antennal cavities separated, the distance between them wider than diameter of cavity, antennae short, can not reach the posterior angle of pronotum, right one with 6 preserved segments, the left with 8 preserved segments, antennomere 1 stout, 2 shorter than 1, with length and width subequal, 3 longer than 2, more slender, 4−8 serrate (
Figs. 9
,
13
).
Pronotum about 1.65 times as wide as long, the widest part of pronotum at its base; anterior pronotal margin straight, posterior margin slightly bisinuate; lateral margins weakly arched at anterior 3/4, then arcuately expanded to posterior margin, basolateral angles acute; integument of pronotum with feeble transverse striae formed by coarse punctates. Scutellum triangular, wider than long, lateral margins arcuately concave, posterior apex acuminate (
Figs. 8
,
12
).
Elytra 3.0 times as long as wide, the widest part at the middle; lateral margins weakly arcuate in anterior 2/3, then somewhat narrowed to obtuse apex; disc with 9 distinct longitudinal striae, the 6th and 7th striae fading out at 1/5 of elytron length from apex; disc with 5 oblong spots: one located at the posterior 2/3 of disc across the forth and the ninth striae; the other four near the inner margin between the first and the sixth striae (
Fig. 8
).
Ventral surface reticulate, finely punctate; prosternal process subparallel, lateral margins slightly converging posteriorly to obtuse apex, process narrower than procoxae; procoxae, mesocoxae rounded, distance between procoxae subequal with distance between mesocoxae; metepisternum quadrate; paracoxal suture straight; metacoxae transverse, short laterally, posterior margin arcuately emarginated; abdominal ventrites with finely, rather dense punctures, which formed into faint rugae, the first ventrite 2.5 times as long as the second one; ventrites 2−3 subequal in length, 4 shorter than 3, 5 broadly rounded at apex, 1.4 times longer than the previous one (
Figs. 9
,
13
).
Legs: pro-, mesotrochanters trapeziform, metatrochanters round triangular; femora subfusiform; tibiae feebly arcuate, slender.
Male genitalia: parameres steeply narrowing to separately acuminate apices; medium lobe robust, with acuminate apex (
Fig. 9
).
Dimensions.
Body length/body width/elytron length, in mm: CNU-COL-NN2010410: 13.6/6.7/10.1.
Remarks.
Among all the species of the fossil subfamily
Parathyreinae
,
Sinoparathyrea robusta
sp. nov.
is the first fossil with the description of the genitalia. It has the similar medium lobe with the species
Agrilus betulanigrae
Macrae, 2003
, whose parameres subquadrately expanded in apical half, wider than the parameres of the species
Sinoparathyres robusta
sp. nov.
.