A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae) Author Duarte, Paulo R. M. Departamento de Fitossanidade, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, Recife, Pernambuco, Brazil Author Grossi, Paschoal C. 0000-0001-6601-5967 Departamento de Fitossanidade, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, Recife, Pernambuco, Brazil & http: // orcid. org / 0000 - 0001 - 6601 - 5967 Author Dupuis, Fabien text Zootaxa 2022 2022-01-25 5093 1 49 66 journal article 2423 10.11646/zootaxa.5093.1.3 701b994e-d6dd-4e39-923d-ad55d113d881 1175-5326 5901043 E90AE625-98EA-41D5-9C8C-CEA4B877E687 Identification key for adults of the Bothynus villiersi species group 1 Apex of the prosternal process conical in both sexes ( Fig. 4C ). If described, female with anterior pronotal border arched ( Fig. 8A )................................................................................................ 2 - Apex of the prosternal process rounded in both sexes ( Fig. 4D ). Anterior pronotal border of female sinuous ( Fig. 8B )......................................................... Bothynus belemensis Duarte, Grossi, & Dupuis , new species 2 Pronotal tubercle of male subtrapezoidal (truncate or bilobed at apex) ( Fig. 4A ). Tergite 8 of female with weakly produced internal plate ( Fig. 8E )................................................... Bothynus alvarengai Martínez, 1983 - Pronotal tubercle of male conical (rounded at apex) ( Fig. 4B ). Tergite 8 of female with strongly produced internal plate ( Fig. 8C–D ).............................................................................................. 3 3 Inner margin of the right paramere bearing a basal process ( Fig. 6D ). Tergite 8 of female with bilobed internal plate ( Fig. 8D )........................................................................ Bothynus horridus Endrödi, 1968 - Inner margin of the right paramere bearing a medial process ( Fig. 6B, H ). Tergite 8 of female parabolic ( Fig. 8C )......... 4 4 Apex of parameres elongate ( Fig. 6B ). Tergite 8 of female with a longitudinal sulcus at the internal plate ( Fig. 8C )................................................................................. Bothynus villiersi Endrödi, 1968 - Apex of parameres shortened ( Fig. 6H ). Female unknow.............. Bothynus spinophallicus (Ratcliffe & Cave, 2021)