<p class = " HeadingRunIn " align = " left "> <strong> Generic classification of the Archiborborinae (Diptera: Sphaeroceridae), with a revision of <em> Antrops </ em> Enderlein, <em> Coloantrops </ em> gen. nov., <em> Maculantrops </ em> gen. nov., <em> Photoantrops </ em> gen. nov., and <em> Poecilantrops </ em> gen. nov. </ strong> </ p> Author KITS, JOEL H. Author MARSHALL, STEPHEN A. text Zootaxa 2013 2013-09-02 3704 1 1 113 http://dx.doi.org/10.11646/zootaxa.3704.1.1 journal article 10.11646/zootaxa.3704.1.1 1175-5326 6507447 341ED5F6-2DCA-4E73-83D6-389DF0DD347F Antrops annulatus (Richards) , comb. nov. ( Figs. 3 , 14 , 21–23 , 195 ) Archiborborus (Procopromyza) annulatus Richards 1963: 232 Description. Head yellow, upper occiput and posterior frons orange to brown, covered with microtomentum except shiny stripes on interfrontal plates outlining frontal triangle. Distance from ocellar bristles to medial ocellus equal to distance between ocelli, ocellar bristles reclinate-divergent. Anterior genal bristle about 0.6X length of vibrissa, no strong subvibrissal bristle. Thorax reddish-brown to dark brown, lateral margins of scutum and pleural sclerites may be mottled with yellow, entirely covered with microtomentum. Scutum with paler dorsocentral stripes, yellow to reddish-brown. Halter whitish, brown at base of knob. Legs yellowish brown to dark brown. Male fore basotarsomere with a small apical spur. Femora 1–3 with a yellow band at two-thirds, otherwise reddish brown to dark brown. Tibia 2 with row of anterodorsal, 1 anteroventral, 1 posteroventral, 4 subapical bristles. Tibia 3 with strong anteroventral, 3 ventroapical bristles. Wing brown. White spots on crossveins r-m and dm-cu, and two on vein R 4+5 distal to r-m crossvein. Abdomen with tergites brown to dark brown, strongly sclerotized, covered with microtomentum. Sternites moderately sclerotized. Tergites 3–5 with a strong bristle in posterolateral corners. Male postabdomen: Sternite 5 with lateral margins rounded, converging, posterior margin with a pair of thin, incurved lobes, anterior apodeme short and broad ( Fig. 23 ). Surstylus very broad, posterior edges nearly meeting medially ( Fig. 22 ). Pregonite distinct, relatively large. Postgonite with lobes widely separated, anterior lobe curved anteriorly, long, rounded, posterior lobe pointed. Basiphallus with broad, very short epiphallus, short, narrow preepiphallus. Ejaculatory apodeme somewhat enlarged. Distiphallus simple, with a smooth, weakly curved dorsal tube ( Fig. 21 ). Female postabdomen: Tergites 6–7 and sternites 6–7 moderately sclerotized, slightly wider than long, entirely covered with microtomentum. Tergites with 3 posterior strips, sternites with 2 posterior strips. Tergite 8 slightly narrower posteriorly, anterior margin shallowly emarginated, posterior two-thirds covered with microtomentum. Epiproct broad, with long anterior arms, posterior portion covered with microtomentum, with 1 pair of setae. Cerci narrow. Sclerites of sternite 8 subrectangular, distal half covered with microtomentum. Hypoproct subrectangular, anterior margin notched, edge lateral to notch shiny. Spermathecae barrel-shaped, about 1.5X longer than wide, with basal and apical invaginations, sclerotized duct short. Type material. Holotype ♂ : CHILE : Biobío : Ñuble , 40 km E of San Carlos , 28 Dec 1950 , Ross & Michelbacher ( CASC ; photo examined) . Paratype ♀ : same data as holotype ( CASC ) . FIGURES 21–24. Antrops male terminalia: A. annulatus : (21) phallus, postgonite, and phallapodeme, lateral view, (22) surstylus, anterior view, (23) sternite 5; A. anovariegatus : (24) phallus, postgonite, and phallapodeme, lateral view. aa, anterior apodeme of sternite 5; bp, basiphallus; dp, distiphallus; dt, dorsal tube of distiphallus; ep, epiphallus; pa, phallapodeme; pep, preepiphallus; prg, pregonite; psgm postgonite. Other material examined. CHILE : Aisén : 15 km S of Las Juntas & 30 km N of Payhuaoi, 100 m , Nothofagus forest, carrion trap , 30 Dec 1984 – 29 Jan 1985 , S. & J. Peck ( 22 ♀ , DEBU ) ; Cisnes Medio , 16 km NW, Rio Grande , 200 m , mature beech forest, carrion trap , 30 Dec 1984 – 28 Jan 1985 , S. & J. Peck ( 2 ♀ , DEBU ) ; Puerto Cisnes , 40 km N, Queulat Pass , 500 m , mossy beech forest, carrion trap , 30 Dec 1984 – 28 Jan 1985 , S. & J. Peck ( 1 ♂ , 1 ♀ , DEBU ) ; Yelcho Chio , Ventisquero Br. , 70 km SE of Chaitén , 200 m , mixed beech forest, carrion trap , 29 Dec 1984 – 29 Jan 1985 , S. & J. Peck ( 2 ♀ , DEBU ) ; Araucanía : Flor del Lago , 15 km NE of Villarrica , 300 m , Nothofagus forest, carrion trap , 14 Dec 1984 – 10 Feb 1985 , S. & J. Peck ( 2 ♂ , DEBU ) ; Lago Tinquilco , 30 km E Pucón , 750 m , yellow pans, 7–10 Dec 2002 , L. Masner ( 5 ♂ , 3 ♀ , DEBU ) ; Malleco , 6 km W Curacautín , 750 m , grazed Nothofagus forest remnant, carrion traps , 12 Dec 1984 – 16 Feb 1985 , S. & J. Peck ( 2 ♀ , DEBU ) ; Nahuelbuta Mts. , 15.6 km W Angol , 600 m , 8 Nov 1966 , M.E. Irwin & E.I. Schlinger ( 1 ♂ , CASC ) ; Princesa , 20 km E of Curacautín , 1500 m , Nothofagus , Malaise , 12 Dec 1984 – 16 Feb 1985 , S. & J. Peck ( 1 ♂ , DEBU ) ; Purén , Contulmo Nat. Mon. , 350 m , mixed evergreen forest, carrion traps , 11 Dec 1984 – 13 Feb 1985 , S. & J. Peck ( 4 ♂ , 8 ♀ , DEBU ) ; Salto de la Princesa , 25 km E Curacautín , 750 m , yellow pans, 7–8 Dec 2002 , L. Masner ( 1 ♀ , DEBU ) ; Termas de Palguín , 33 km E Pucón , 680 m , along spring in woods, 6 Nov 1989 , S.A. Marshall ( 1 ♀ , DEBU ) ; Biobío : Alto Tregualemu , ca. 20 km SE Chovellen , 500 m , 26–27 Jan 1979 , D. & M. Davis , B. Akerbergs ( 1 ♀ , USNM ) ; Concepción , 6km S., San Pedro, 360 m , 12 Dec 1982 – 2 Jan 1983 , A. Newton & M. Thayer ( 14 ♂ , 7 ♀ , DEBU , CNCI ) ; as above but Pinus sp. forest, trap site 648 ( 2 ♂ , DEBU ) ; Laraquete , yellow pans, 10–25 Jan 1993 , P. Salinas ( 24 ♂ , 11 ♀ , DEBU ) ; Santa Bárbara , 25 km E, 350 m , 24 Jan 1978 , W.N. Mathis ( 1 ♀ , USNM ) ; near coastal stream, 17.5 km S Curanipe , 50 m , 25 Jan 1979 , D. & M. Davis , B. Akerbergs ( 1 ♀ , USNM ) ; Los Lagos : Alerce Andino Natl. Pk. , trail to Laguna Fría , 400 m , dung & carrion pans, 1 Dec 2008 , Kits & Marshall ( 3 ♂ , 3 ♀ , DEBU ) ; Anticura , 4 km E, 430 m , 19–26 Dec 1982 , Newton & Thayer ( 1 ♀ , DEBU ) ; Antillanca , 2.5 km W, 950 m , dung trap , 18 Dec 1977 , [no collector] ( 6 ♂ , 2 ♀ , USNM ) ; Bahía Mansa , 3 km S Maicolpue , 200 m , mixed forest, carrion trap , 21 Dec 1984 – 3 Feb 1985 , S. & J. Peck ( 1 ♂ , 2 ♀ , DEBU ) ; Chepu , Chiloé , 40 ft , Nothofagus forest, 3 Oct 1958 , Royal Society Chilean Exp. ( 2 ♂ , BMNH ) ; Chiloé I. , 11 km NW of Castro , 200 m , ravine, 2nd growth forest, carrion trap , 27 Dec 1984 – 1 Feb 1985 , S. & J. Peck ( 4 ♂ , 4 ♀ , DEBU ) ; Chiloé I. , Chiloé Natl. Pk. , 0 m, yellow pans, 11–12 Dec 2002 , L. Masner ( 4 ♂ , 1 ♀ , DEBU ) ; Chiloé I. , Lago Huinchillo , 10 m , yellow pans, 11–12 Dec 2002 , L. Masner ( 2 ♂ , 5 ♀ , DEBU ) ; Chiloé I. , Miraflores , Lago Tepuhuico , 15 m , shoreline forest, yellow pans, 13–15 Feb 2005 , [no collector] ( 2 ♂ , 2 ♀ , DEBU ) ; Chiloé I. , Miraflores , Punto Río Negro , 50 m , creek, yellow pans, 14–15 Feb 2005 , L. Masner ( 1 ♂ , 1 ♀ , DEBU ) ; P.N. Puyehue , 1370 m , 20–25 Dec 1982 , A. Newton & M. Thayer ( 1 ♂ , DEBU ) ; Parque Nac. Puyehue , 1600 m , 18–24 Dec 1982 , A. Newton & M. Thayer ( 1 ♂ , DEBU ) ; as above but 690 m ( 1 ♂ , DEBU ) ; as above but 965 m , 18–25 Dec 1982 ( 1 ♀ , DEBU ) ; Parque Nac. Puyehue , 4.1 km E Anticura , 430 m , Malaise trap , 19–25 Dec 1982 , N.E. Woodley ( 1 ♂ , USNM ) ; Pucatrihue , 27–30 Jan 1978 , W.N. Mathis ( 1 ♀ , USNM ) ; Puyehue Nat. Park , Anticura , 250 m , Nothofagus forest, 13 Feb 1988 , L. Masner , Chile Exp. ( 1 ♂ , CNCI ) ; Puyehue Natl. Pk. , 1600 m , 18–24 Dec 1982 , Newton & Thayer ( 1 ♀ , DEBU ) ; Puyehue Natl. Pk. , Anticura , 375 m , 3–4 Dec 2008 , Kits & Marshall ( 1 ♀ , DEBU ) ; Puyehue Natl. Pk. , Anticura , Repucura forest trail, 500 m , carrion trap , 19 Dec 1984 – 6 Feb 1985 , S. & J. Peck ( 2 ♀ , DEBU ) ; Puyehue Natl. Pk. , Anticura , trail to Mirador El Puma , 400 m , carrion pans, 3–5 Dec 2008 , Kits & Marshall ( 1 ♂ , 2 ♀ , DEBU ) ; as above but dung pans ( 2 ♂ , 6 ♀ , DEBU ) ; Río San Pedro , 24 km NE of Los Lagos , river bank slope forest, yellow pans, 13–18 Nov 2000 , L. Packer ( 2 ♂ , 12 ♀ , DEBU ) ; Termas de Puyehue , 8 km E, woods, dung trap , 9 Dec 1977 , [no collector] ( 2 ♀ , USNM ) ; Vicente Pérez Rosales Natl. Pk., Volcan Osorno rd., 300 m , carrion pans, 2 Dec 2008 , Kits & Marshall ( 4 ♂ , 1 ♀ , DEBU ) ; as above but dung pans ( 1 ♂ , DEBU ) ; Los Ríos : Panguipulli , 26 km SE, 300 m , carrion trap , Nothofagus remnant, 16 Dec 1984 – 11 Feb 1985 , S. & J. Peck ( 3 ♀ , DEBU ) ; Puerto Fuy , 800 m , Nothofagus , 16 Dec 1984 – 11 Feb 1985 , S. & J. Peck ( 1 ♀ , DEBU ) ; Magallanes : Lago del Toro , 3 km E, 300 m , dung trap , 7 Dec 1977 , [no collector] ( 7 ♂ , 3 ♀ , USNM ) ; Maule : Altos del Lircay Natl. Res. , 1256 m , 29 Nov 2006 , S.A. Marshall ( 2 ♂ , 4 ♀ , DEBU ) ; as above but ca. 3000 m ( 5 ♂ , 2 ♀ , DEBU ) ; as above but streamside pans ( 9 ♂ , 4 ♀ , DEBU ) ; Constitucíon , 16 Dec 1976 , Gurney & Barria ( 1 ♀ , USNM ) ; Pelluhue , along Río Curanilahue , 26 Nov 2006 , S.A. Marshall ( 2 ♂ , 1 ♀ , DEBU ) ; Putú , ca. 15 km N Constitución , along stream, 26 Nov 2006 , S.A. Marshall ( 4 ♂ , 6 ♀ , DEBU ) ; Valparaíso : La Campana Nat. Pk. , 900 m , hygrophilous forest, FIT, 2 Dec 1984 – 21 Feb 1985 , S. & J. Peck ( 3 ♀ , DEBU ) ; Zapallar , Quebrada El Tigre , above tennis club, 100 m , 9 Dec 2008 , Kits & Marshall ( 1 ♂ , 1 ♀ , DEBU ) . Comments. A. annutalus is quite distinctive within the genus, particularly the banded legs and striped scutum. Only A. vittatus sp. nov. is likely to be confused with this species, but is readily distinguished by the colour of the femora and the leg chaetotaxy.