Review of the mirine plant bug genus Eurystylus Stål from Japan and Taiwan (Hemiptera: Heteroptera: Miridae: Mirinae), with descriptions of two new species, a new synonymy and a new combination
Author
Yasunaga, Tomohide
Author
Nakatani, Yukinobu
Author
Chérot, Frédéric
text
Zootaxa
2017
4227
3
301
324
journal article
37230
10.11646/zootaxa.4227.3.1
d71680f0-0069-4290-a53e-9272043455fc
1175-5326
268312
62ABB516-62B1-46AF-A235-3AD772358A8A
Genus
Eurystylopsis
Poppius
Eurystylopsis
Poppius, 1911
: 18
(n. gen.),
type
species:
Eurystylopsis longipennis
Poppius, 1911
: 19
(
INDIA
:
Darjeeling
), original designation;
Schuh
, 1995
: 765
(cat.);
Kerzhner
&
Josifov
, 1999
: 98
(cat.);
Zheng
et al.
, 2004
: 264
(diag., key to
Chinese
spp.);
Schuh
(
2002–2014
) online catalog.
Eurystylomorpha
Poppius, 1915
: 16
(n. gen.),
type
species:
Eurystylomorpha crassicornis
Poppius, 1915
: 17
(
TAIWAN
:
Fuhosho
), original designation;
Schuh
, 1995
: 765
(cat.);
Kerzhner
&
Josifov
, 1999
: 98
(cat.);
Zheng
et al.
, 2004
: 262
(diag.);
Schuh
(
2002–2014
) online catalog.
n. syn.
Diagnosis.
Eurystylopsis
is distinguished from other mirine genera by a combination of the following characters: Body elongate ovoid, not much tumid nor boxlike; basic coloration dark to reddish brown; general coloration and size more or less sexually dimorphic (as in
E. clavicornis
,
Figs. 8
H–I, 9A–B); antennal segment II clavate, apparently longer than basal width of pronotum, with its apical part more than twice as thick as base; segments III and IV short, filiform; pronotum often with a few dark stripes in female (9B–D); collar narrow, about as thick as base of antennal segment II; scutellum moderately (as in
E. clavicornis
,
Fig. 9
A–B) or sometimes strongly (as in
E. harmandi
,
Fig. 9
B–E) swollen; legs long; left paramere C-shaped, weakly constricted subapically (
Fig. 10
A); right paramere short, straight (
Fig. 10
B); endosoma with notched sclerites and rather small, thick-rimmed secondary gonopore (
Fig. 10
C); female bursa copulatrix with thick-rimmed, large sclerotized ring (10D); and posterior wall with developed, T- or Y-shaped dorsal structure and bilobate interramal lobe (
Fig. 10
E).
Distribution.
China
,
India
,
Nepal
and
Taiwan
.
Discussion.
This genus is considered to be the most closely related taxon to
Eurystylus
. Two species were found to be associated with inflorescences of chestnut,
Castanea
sp. and orange,
Citrus
sp. (
Eurystylopsis clavicornis
) and an undetermined broadleaf (
Ep. hermandi
,
Fig. 9
C–D) in
Nepal
. These observations may imply a relationship
between
Eurystylus
and
Eurystylopsis
.
Poppius (1915)
described a new genus
Eurystylomorpha
to accommodate a single species,
Em. crassicornis
(
Fig. 8
D) from
Taiwan
, comparing it with
Eurystylus
only. We find it mysterious that he (loc. cit.) did not mention his own genus
Eurystylopsis
proposed for two Himalayan species just four years prior (
Poppius, 1911
). We imagine that Poppius did not included
Eurystylopsis
because the geographical distribution of the genera in
India
and
Taiwan
are remotely disjunct. However, four additional species were subsequently described from continental
China
(
Zheng & Chen, 1991
;
Zheng
et al.
, 2004
). We have also found
E. clavicornis
distributed in
Nepal
and
Thailand
, which could connect the distribution range for the genus. Both nominal genera share numerous character states, except for the carina on vertex (absent in
Eurystylopsis
; thin but complete in
Eurystylomorpha
), the overlapping of mesoscutum, and the coloration of dorsal pilosity. These minor differences are now considered insufficient to recognize them as two independent genera. We therefore suggest a new subjective synonymy,
Eurystylopsis
Poppius, 1911
=
Eurystylomorpha
Poppius, 1915
. Accordingly,
Eurystylopsis
is now known widely from the Oriental to eastern Palearctic regions.