Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species Author Galindo, Lee Ann B84DC387-F1A5-4FE4-80F2-5C93E41CEC15 Muséum national d’Histoire naturelle, Département Systématique et Evolution, ISyEB Institut (UMR 7205 CNRS / UPMC / MNHN / EPHE), 43, Rue Cuvier, 75231 Paris, France. & urn: lsid: zoobank. org: author: B 84 DC 387 - F 1 A 5 - 4 FE 4 - 80 F 2 - 5 C 93 E 41 CEC 15 & Corresponding author: galindo @ mnhn. fr galindo@mnhn.fr Author Kool, Hugo H. 5E718E5A-85C8-404C-84DC-6E53FD1D61D6 Associate Mollusca Collection, Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. & Email: hugo. kool @ naturalis. nl & urn: lsid: zoobank. org: author: 5 E 718 E 5 A- 85 C 8 - 404 C- 84 DC- 6 E 53 FD 1 D 61 D 6 hugo.kool@naturalis.nl Author Dekker, Henk DA6A1E69-F70A-42CC-A702-FE0EC80D77FA Associate Mollusca Collection, Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. & Email: h-dekker @ quicknet. nl & urn: lsid: zoobank. org: author: DA 6 A 1 E 69 - F 70 A- 42 CC-A 702 - FE 0 EC 80 D 77 FA h-dekker@quicknet.nl text European Journal of Taxonomy 2017 2017-02-10 275 1 43 journal article 22200 10.5852/ejt.2017.275 80a9157d-3178-4f1b-9c98-436636c4d4d0 2118-9773 3824544 FC663FAD-BCCB-4423-8952-87E93B14DEEA Reticunassa thailandensis sp. nov. urn:lsid:zoobank.org:act: E0731396-9A77-4C1C-99F7-24B72E5B55DD Fig. 8 M–P, 12; Table 2 Etymology This species is named after Thailand , the type locality. Type material Holotype THAILAND : SE coast, Gulf of Thailand , Pattani Province , Ko Lopi , 06°40.313′ N , 101°43.852′ E , scuba diving, 3–15 m , leg. R.G. Moolenbeek and H. Dekker , 2003 ( ZMA.Moll.4.09.047 , length 7.35 mm , width 3.8 mm ). Paratypes THAILAND : same collection data as for holotype ( ZMA .Moll.4.09.048, 222 spm; 5 spm; MNHN IM- 2000-32481, 10 spm; GH , 2 spm; HD 24145, 6 spm; HK 139.01, 6 spm); Gulf of Thailand , Rayong Province , Talengam, near Suan Son, E of Ban Phe beach, leg. O.K. McCausland ( HK 139.03, 1 spm). Other material examined THAILAND : Gulf of Thailand: type locality (4 spm); Ban Pak Nam Sakom, local fishery, 06°57′ N , 100°49.36′ E ( ZMA .Moll.4.09.062, 3 spm; HD 1789, 6 spm); Ban Pak Nam Sakom, near Leela Resort, 06°57.4′ N , 100°51.6′ E , muddy pools ( ZMA , 3 spm; HD 1190, 6 spm); Ban Talokapo, 06°54.0′ N , 101°22.4′ E , local fishery ( HD 2146, 3 spm); Trat Province , Ko Chang, beach ( HD 2793, 1 spm); Ko Samui, NE side of Chaweng Beach, 09°32.6′ N , 100°05.7′ E , in seagrass ( HD 9572, 5 spm); Ko Samui, Ban Bangkao, 09°25. 2′ N , 99°59.7′ E , sand flat ( HD 9574, 1 spm); Ko Kham, 06°58.4′ N , 100°51.6′ E , scuba diving, 3–5 m ( ZMA .Moll.093866, 14 spm); Pattani , in shell grit ( ZMA .Moll.098947, 2 spm); off Pattaya, Ko Lan, scuba diving, reef, 3–6 m , leg. R.G. Moolenbeek ( ZMA .Moll.4.09.063, 12 spm). SINGAPORE : Bedok River area of East Coast Park, beached, leg. Chan Sow Yan ( HK 139.04, 6 spm). MALAYSIA : E coast, South China Sea, 02°38′11″ N , 105°13′30″ E , 30–35 m , leg. H. Morrison ( GH , 2 spm; HK 139.02, 3 spm); South China Sea, on sand near wreck, 0°45′15″ N , 105°36′11″ E , leg. H. Morrison ( GH , 1 spm). INDONESIA : Laboratorium Onderzoek der Zee, 1938, stn Bat38/07a, Bay of Jakarta ( ZMA . Moll.096217, 2 spm). – KEPULAUAN SERIBU 2005, stn Ser05/03, Java , Bay of Jakarta , Damar Kecil, 5°59′00″ S , 106°50′43″ E , scuba diving, 5–15 m ( ZMA .Moll.175169, 7 spm); stn ser05/07, Java , Bidadari, scuba diving, 6°01′55″ S , 106°44′47″ E ( ZMA .Moll.175470, 12 spm); stn ser05/08, Java , Ayer Besar, 6°00′06″ S , 106°46′48″ E ( ZMA .Moll.154506, 1 spm); stn ser05/09, Java , Ubi Besar, scuba diving, 5°59′55″ S , 106°44′23″ E ( ZMA .Moll.175172, 8 spm); stn ser05/10, Java , Kelor, scuba diving, 5°51′00″ S , 106°35′05″ E , 5–35 m ( ZMA .Moll.175173, 3 spm). – Java , W coast, Tjilintjing ( ZMA .Moll.096215, ex coll. Butot 12405, 1 spm); Flores, E coast, near Larantuka, between Weri and Waibalun, leg. Fr.M. Vianney ( ZMA .Moll.096211, 1 spm); Kalimantan , South China Sea, Pejantan I., 0°30′48.8″ N , 107°13′24″ E , 10–20 m , leg. H. Morrison ( GH , 5 spm); NETHERLANDS BERAU 2003, stn ber03/38, Berau Is, Maratua, 10–25 m ( ZMA .Moll.175049, 1 spm). PHILIPPINES : Palawan , Balabac I. ( RMNH , 5 spm; 5 spm; HD 24144, 20 spm; GH , 6 spm; HK 139.05, 140 spm; coll. L. van Gemert, Zeist, the Netherlands , 20 spm; AMD ,>300 spm); Olango I., Ziczac Point, 10–20 m , 2010 ( HD 33413, 1 spm); Mindanao, Surigao del Norte, Basul I., 80–90 m , 2013 ( HD 33414, 1 spm). Description Holotype PROTOCONCH. Bulbous, flattened, paucispiral, 1.3 whorls, whitish, part of body whorl with growth lines. SHELL. Teleoconch consisting of 4.5 impressed whorls.Axial ribs pronounced, rounded, 14 on penultimate and 13 on body whorl. Varix strong and broad. SPIRAL CORDS. Continuous, narrow, thread-like spiral cords, increasing in height where they cross over axial ribs; 6 on penultimate and approximately 9 on body whorl. Spiral cord on periphery, the two above it, and the three at the base of body whorl more distinct; intercordal distances unequal. INTERCORDAL SCULPTURE. Fine, distinct spiral striae, number depending on distance between spiral cords. APERTURE. Oval, outer lip with 8 unequal denticles, one near periphery strongest. Columellar callus well bordered, slightly extending over body whorl; entire lip with short lirae. Parietal denticle strong. OPERCULUM. Yellowish, finely serrated. SIPHONAL CANAL. Narrow; fasciole strong. Siphonal area with approximately 4 cords; parietal denticle strong, anal canal wide. COLOR. Yellowish-orange, penultimate whorl with two, body whorl with three orange-brown bands. ADULT SIZE. 5.5–9.1 mm , usually 7–8 mm . One of the smaller species of the group. Remarks The color is variable. Well-preserved specimens are cream or orange to light brown, either unicolor or with several darker-colored spiral cords. Occasionally remnants of bands are only present on the varix and on the fasciole. The main difference between Reticunassa intrudens sp. nov and R. thailandensis sp. nov. is to be found in the protoconch morphology: the former has a protoconch with axial striae, in the latter it is partly smooth. Both are paucispiral and bulbous, and lack the microscopic pustules found in the other species described in this paper. R. paupera ( Gould, 1850 ) , R. visayaensis sp. nov. and R. poppeorum sp. nov. have a multispiral protoconch. R. thailandensis sp. nov. has more well developed spiral cords than the other species described here. Furthermore, in some specimens the 2–3 most anterior spiral cords are more strongly developed. In general, the distribution of the species, as shown on Fig. 12 , might help with the identification. Habitat This species lives in sand and between dead corals, in depths of 1–90 m , usually found between 15 and 20 m . Distribution Thailand , Singapore , Malaysia , Indonesia , and the Philippines , where it is common at Balabac I., but apparently rare in the central Visayas ( Fig. 12 ).