The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea
Author
Harzhauser, Mathias
0000-0002-4471-6655
Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655
mathias.harzhauser@nhm-wien.ac.at
Author
Landau, Bernard M.
0000-0002-4471-6655
Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655
mathias.harzhauser@nhm-wien.ac.at
text
Zootaxa
2024
2024-03-19
5427
1
1
110
http://dx.doi.org/10.11646/zootaxa.5427.1.1
journal article
10.11646/zootaxa.5427.1.1
1175-5334
10841061
923206B0-E8C5-4FD5-B882-55009ABB0282
Family
Tudiclidae
Cossmann, 1901
Revised diagnosis.
“
Shell medium-sized to medium-large, from 10 to about
100 mm
in adults, with very short to very long siphonal canal, sometimes twisted or strongly inclined abaxially. Protoconch paucispiral, usually large to very large and bulbous. Axial sculpture usually present at least on adapical whorls, of rounded axial ribs or knobs varying in strength, rarely absent. Spiral sculpture completely absent, or represented by striae or cords of variable strength and density. Outer aperture lip smooth or lirate inside, inner lip calloused, sometimes bearing a parietal knob
.” (
Kantor
et al.
2022: 832
).
Discussion.
The family was revised by Kantor
et al.
(2021) based on molecular data. The living
Euthria
J.E.
Gray, 1850
and
Tudicla
Röding, 1798
were placed in this family by Kantor
et al.
(2021). In addition, the extinct
Euthriofusus
Cossmann, 1901
, was moved into
Tudiclidae
by
Lozouet (2021: 33)
based on conchological similarities with the
Tudiclidae
genus
Afer
Conrad, 1858
. The family can be traced back to the Late Oligocene with species, such as
Euthriofusus peyreirensis
Peyrot, 1928
, from the northeastern Atlantic. This occurrence might hint at an Eocene/Early Oligocene origin in the tropical Eastern Atlantic. Cretaceous genera, listed in
Tudiclidae
by
Saul (1988)
and
Harasewych (2018)
belong to extinct families, such as Pyropsinae
Stephenson, 1941
(within
Pholidotomidae
Cossmann, 1896
),
Perissityidae
Popenoe & Saul, 1987
and
Johnwyattidae
Serna, 1979
(see
Bandel 1993
;
Snyder 2003
).
Genus
Tudicla
Röding, 1798
Type
species.
Murex spirillus
Linné, 1767
; subsequent designation by
Angas (1878: 611)
. Present-day, Indo-West Pacific.
Revised diagnosis.
“[…]
rounded body whorl, distinctive inductura, a long, well-demarcated siphonal canal, as well as a sharply defined siphonal fold
[…]” (
Harasewych 2018: 36
).
Discussion.
Tudicla
is characterized by an extraordinarily long siphonal canal, a blunt columellar fold and prominent parietal fold. A large, mammillate protoconch is also typical for
Tudicla
and some other
Tudiclidae
such as
Afer
Conrad, 1858
(
Fraussen 2008
;
Harasewych 2018
). First records of this genus probably date back to the Late Oligocene of
Japan
(
e.g
.,
Tudicla japonica
Takeda, 1953
,
Tudicla ishii
Matsui, 1959
, see
Takeda 1953
: pl. 2, figs 13–15;
Matsui 1959
: pl. 2, fig. 4), but the poor preservation does not allow a clear identification. The oldest undoubted occurrences are Early Miocene records of
Tudicla rusticula
(de
Basterot, 1825
)
from the Circum-Mediterranean Region. During the Middle Miocene,
Tudicla hoernesi
(
Stur, 1870
)
represents a second European
Tudicla
species.
Additional Middle Miocene species might be represented by
Tudicla angulata
Tate, 1888
,
T. costata
Tate, 1888
and
T. turbinata
Tate, 1888
from southern
Australia
(age according to
Squires 2011
). These species will need confirmation, because the illustrations in
Tate (1888)
do not show any of the important apertural features.
Tudicla
was also reported from Cretaceous, Paleocene and Eocene strata (
e.g
.,
Cox 1925
;
Traub 1979
;
Abdel-Gawad 1986
) but in our opinion ‘
Tudicla’ krenkeli
Cox, 1925
, from the Maastrichtian of
Mozambique
, and several species described by
Abdel-Gawad (1986)
from the Maastrichtian of
Poland
, belong in the Pyropsinae
Stephenson, 1941
. Similarly,
Squires (2011: 1204)
rejected Paleocene and Eocene occurrences and excluded these from
Tudicla
. For synonyms of
Tudicla
see
Harasewych (2018: 36)
.