<document id="1485529AFEA5D3688070C33287D64F02" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFB6844F8CFBFEABDFE8FDC1" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes panamensis" docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="44" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="95E83531A4B3208F0092F4033F8ADA69">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:namePart id="69EC35E610256635B66E5B2F9900FB1E">Kluge, Nikita J.</mods:namePart>
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<mods:date id="CAF0E279A2C7D6A15EBB366A15500B8A">2020</mods:date>
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<mods:number id="BF60136BF61C5E52B7A1138BDF849BD9">4756</mods:number>
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Group 
<taxonomicName id="4C3E4D33FFB6844F8C9DFEABDABEFEC5" box="[241,384,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B94AEAA2FFB6844F8C9DFEABDABEFEC5" bold="true" box="[241,384,333,359]" italics="true" pageId="43" pageNumber="44">panamensis</emphasis>
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<subSubSection id="C324653BFFB6844F8CFBFE72D948FDE2" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8B8136B0FFB6844F8CFBFE72D948FDE2" blockId="43.[151,1436,404,611]" pageId="43" pageNumber="44">
<emphasis id="B94AEAA2FFB6844F8CFBFE72DA28FE0D" bold="true" box="[151,278,404,431]" pageId="43" pageNumber="44">Definition.</emphasis>
Penis lobes moderately long and moderately widened apically, with narrow lateral pouches; telopenes in form of spear-like rolls, attached on dorsal side and directed medially-dorsally (
<figureCitation id="13052A35FFB6844F8820FE5FDFD2FE71" box="[1100,1260,440,467]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="43" pageNumber="44">Figs 143–145</figureCitation>
, 
<figureCitation id="13052A35FFB6844F8896FE5FDE61FE71" box="[1274,1375,441,467]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="43" pageNumber="44">169, 171</figureCitation>
) (the same in many other species). In imago and subimago of both sexes, femora with one multicolored apical band (
<figureCitation id="13052A35FFB6844F8906FE3BDBC0FDBE" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="43" pageNumber="44">Figs 160–161</figureCitation>
); in male, abdominal terga II–VI whitish, terga VII–X orange (
<figureCitation id="13052A35FFB6844F8FD7FDE7DF1EFDB9" box="[955,1056,513,539]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="43" pageNumber="44">Figs 136</figureCitation>
, 
<figureCitation id="13052A35FFB6844F8840FDE7DF69FDBE" box="[1068,1111,513,540]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="43" pageNumber="44">166</figureCitation>
) (the same coloration of legs and male abdomen in many other species).
</paragraph>
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<emphasis id="B94AEAA2FFB6844F8CABFDAEDA59FDC1" bold="true" box="[199,359,584,611]" pageId="43" pageNumber="44">Composition.</emphasis>
Among species described here, 
<taxonomicName id="4C3E4D33FFB6844F8EA3FDAFD841FDC1" box="[719,895,585,611]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB6844F8EA3FDAFD841FDC1" box="[719,895,585,611]" italics="true" pageId="43" pageNumber="44">Th. panamensis</emphasis>
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<emphasis id="B94AEAA2FFB6844F8FEAFDAFD8F8FDC1" bold="true" box="[902,966,585,611]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="A27957D9FFB6844F8FEAFDAFD8F8FDC1" box="[902,966,585,611]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C3E4D33FFB6844F8F90FDAFDFB1FDC1" box="[1020,1167,585,611]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
<emphasis id="B94AEAA2FFB6844F8F90FDAFDFB1FDC1" box="[1020,1167,585,611]" italics="true" pageId="43" pageNumber="44">Th. viviparus</emphasis>
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<emphasis id="B94AEAA2FFB6844F88FAFDAFDFE8FDC1" bold="true" box="[1174,1238,585,611]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="A27957D9FFB6844F88FAFDAFDFE8FDC1" box="[1174,1238,585,611]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
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